List of rodents

five rodent images
Clockwise from top-left: capybara (Hydrochoerus hydrochaeris), South African springhare (Pedetes capensis), golden-mantled ground squirrel (Callospermophilus lateralis), house mouse (Mus musculus), North American beaver (Castor canadensis)
Map of the world, with most of the world outside of Antarctica and the furthest northern arctic regions shaded blue
Worldwide distribution of rodent species

Rodentia is an order of placental mammals. Members of this order are called rodents. The order comprises 2,360 extant species, which are grouped into 511 genera. The largest order of mammals, rodents comprise about 40% of all mammal species worldwide. They are native to all major land masses except for Antarctica, and can be found worldwide. They live in a variety of habitats, particularly forests, grasslands, and shrublands, but also savannas, wetlands, deserts, and rocky areas. Most rodents are small animals with robust bodies, short limbs, and long tails. They range in length from the African pygmy mouse, at 4 cm (2 in) plus a 2 cm (1 in) tail, to the capybara, at 134 cm (53 in) plus a 2 cm (1 in) tail.

Rodentia is divided into five suborders: Anomaluromorpha, Castorimorpha, Hystricomorpha, Myomorpha, and Sciuromorpha. The suborders are further subdivided into clades and families. Anomaluromorpha contains two families, the scaly-tailed squirrels and the springhares. Castorimorpha contains three families and includes beavers, gophers, kangaroo rats, and pocket mice. Hystricomorpha contains 18 families and includes mole-rats, porcupines, cavies, hutias, tuco-tucos, and Neotropical spiny rats. Myomorpha contains nine families and includes jerboas, hamsters, voles, and a wide variety of rats and mice. Sciuromorpha contains three families and includes squirrels, chipmunks, and dormice.[1][2] The exact organization of the species is not fixed, with many recent proposals made based on molecular phylogenetic analysis. In addition to the extant rodents, 38 species have been driven extinct since 1500 CE.

Conventions

The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Range maps are provided wherever possible; if a range map is not available, a description of the collective range of species in that genera is provided. Ranges are based on the International Union for Conservation of Nature (IUCN) Red List of Threatened Species unless otherwise noted. All extinct genera or species listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "".

Classification

The order Rodentia consists of 2,360 extant species belonging to 511 genera. This does not include hybrid species or extinct prehistoric species. Modern molecular studies indicate that the 511 genera can be grouped into 35 families; these families are divided between 5 named suborders and are grouped in those suborders into named clades, and some of these families are subdivided into named subfamilies. An additional 38 species have been driven extinct since 1500 CE: 8 in the family Capromyidae, 6 in Echimyidae, 11 in Cricetidae, and 13 in Muridae.

Suborder Anomaluromorpha

  • Family Anomaluridae (scaly-tailed squirrels): 3 genera, 7 species
  • Family Pedetidae (springhares): 1 genus, 2 species

Suborder Castorimorpha

  • Superfamily Castoroidea
  • Superfamily Geomyoidea
    • Family Geomyidae (gophers): 7 genera, 36 species
    • Family Heteromyidae (kangaroo rats and picket mice)
      • Subfamily Dipodomyinae (kangaroo rats, and kangaroo mice): 2 genera, 22 species
      • Subfamily Heteromyinae (spiny pocket mice): 1 genus, 14 species
      • Subfamily Perognathinae (pocket mice): 2 genera, 27 species

Suborder Hystricomorpha

  • Parvorder Caviomorpha
    • Superfamily Cavioidea
      • Family Caviidae (cavies)
        • Subfamily Caviinae (guinea pigs and cavies): 3 genera, 15 species
        • Subfamily Dolichotinae (maras): 1 genus, 2 species
        • Subfamily Hydrochoerinae (capybaras and rock cavies): 2 genera, 4 species
      • Family Cuniculidae (pacas): 1 genus, 2 species
      • Family Dasyproctidae (agoutis and achouchis): 2 genera, 15 species
      • Family Erethizontidae (New World porcupines)
        • Subfamily Chaetomyinae (bristle-spined rat): 1 genus, 1 species
        • Subfamily Erethizontinae (prehensile-tailed porcupines): 2 genera, 17 species
    • Superfamily Chinchilloidea
      • Family Dinomyidae (pacarana): 1 genus, 1 species
      • Family Chinchillidae (chinchillas and viscachas): 2 genera, 7 species (1 extinct)
    • Superfamily Octodontoidea
      • Family Abrocomidae (chinchilla rats): 2 genera, 10 species
      • Family Capromyidae (hutias)
        • Subfamily Capromyinae (hutias): 4 genera, 11 species (2 extinct)
        • Subfamily Hexolobodontinae  (imposter hutia): 1 genus, 1 species (1 extinct)
        • Subfamily Isolobodontinae  (laminar-toothed hutias): 1 genus, 2 species (2 extinct)
        • Subfamily Plagiodontinae (Hispaniolan hutias): 3 genera, 4 species (3 extinct)
      • Family Ctenomyidae (tuco-tucos): 1 genus, 59 species
      • Family Echimyidae (Neotropical spiny rats)
        • Subfamily Carterodontinae (Owl's spiny rat): 1 genus, 1 species
        • Subfamily Echimyinae (tree-rats, bamboo rats, and brush-tailed rats): 18 genera, 75 species
        • Subfamily Euryzygomatomyinae (Atlantic spiny rats): 3 genera, 12 species
        • Subfamily Heteropsomyinae (cave rats): 3 genera, 6 species (6 extinct)
      • Family Octodontidae (degus, rock rats, and viscacha rats): 7 genera, 14 species
  • Superfamily Ctenodactyloidea
  • Family Hystricidae (Old World porcupines): 3 genera, 11 species
  • Parvorder Phiomorpha
    • Family Bathyergidae (mole-rats): 5 genera, 21 species
    • Family Heterocephalidae (naked mole-rat): 1 genus, 1 species
    • Family Petromuridae (dassie rat): 1 genus, 1 species
    • Family Thryonomyidae (cane rats): 1 genus, 2 species

Suborder Myomorpha

  • Superfamily Dipodoidea
    • Family Dipodidae (jerboas)
      • Subfamily Allactaginae (jerboas): 4 genera, 16 species
      • Subfamily Cardiocraniinae (pygmy jerboas): 3 genera, 7 species
      • Subfamily Dipodinae (three-toed jerboas): 5 genera, 9 species
      • Subfamily Euchoreutinae (long-eared jerboa): 1 genus, 1 species
    • Family Sminthidae (birch mice): 1 genus, 16 species
    • Family Zapodidae (jumping mice): 3 genera, 5 species
  • Superfamily Muroidea
    • Family Calomyscidae (brush-tailed mice): 1 genus, 8 species
    • Family Cricetidae (New World rats and mice)
      • Subfamily Arvicolinae (voles, lemmings, and muskrats): 30 genera, 158 species
      • Subfamily Cricetinae (hamsters): 7 genera, 18 species
      • Subfamily Neotominae (deer mice, packrats, and grasshopper mice): 16 genera, 124 species (1 extinct)
      • Subfamily Sigmodontinae (New World rats and mice): 87 genera, 395 species (10 extinct)
      • Subfamily Tylomyinae (vesper rats and climbing rats): 4 genera, 10 species
    • Family Muridae (Old World rats and mice)
      • Subfamily Deomyinae (spiny mice and brush-furred rats): 4 genera, 55 species
      • Subfamily Gerbillinae (gerbils, jirds, and sand rats): 14 genera, 102 species
      • Subfamily Leimacomyinae (Togo mouse): 1 genus, 1 species
      • Subfamily Lophiomyinae (maned rat): 1 genus, 1 species
      • Subfamily Murinae (Old World mice and rats): 136 genera, 655 species (13 extinct)
    • Family Nesomyidae (African and Malagasy rats and mice)
      • Subfamily Cricetomyinae (hamster-rats and pouched rats): 3 genera, 8 species
      • Subfamily Delanymyinae (Delany's mouse): 1 genus, 1 species
      • Subfamily Dendromurinae (climbing mice and fat mice): 6 genera, 26 species
      • Subfamily Mystromyinae (white-tailed rat): 1 genus, 1 species
      • Subfamily Nesomyinae (Malagasy rodents): 9 genera, 27 species
      • Subfamily Petromyscinae (rock mice): 1 genus, 4 species
    • Family Platacanthomyidae (Oriental dormice): 2 genera, 2 species
    • Family Spalacidae (mole-rats)
      • Subfamily Myospalacinae (zokors): 2 genera, 6 species
      • Subfamily Rhizomyinae (bamboo rats and mole-rats): 3 genera, 6 species
      • Subfamily Spalacinae (blind mole-rats): 2 genera, 11 species

Suborder Sciuromorpha

  • Family Aplodontiidae (mountain beaver): 1 genus, 1 species
  • Family Gliridae (dormice)
    • Subfamily Glirinae (Japanese dormice and edible dormice): 2 genera, 2 species
    • Subfamily Graphiurinae (African dormice): 1 genus, 15 species
    • Subfamily Leithiinae (dormice): 6 genera, 12 species
  • Family Sciuridae (squirrels, chipmunks, and marmots)
    • Subfamily Callosciurinae (Asian squirrels): 14 genera, 67 species
    • Subfamily Ratufinae (giant squirrels): 1 genus, 4 species
    • Subfamily Sciurillinae (Neotropical pygmy squirrel): 1 genus, 1 species
    • Subfamily Sciurinae (tree squirrels and flying squirrels): 20 genera, 2 species
    • Subfamily Xerinae (ground squirrels, chipmunks, and prairie dogs): 24 genera, 2 species
Rodentia[3]

Rodents

The following classification is based on the taxonomy described by Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[1][2]

Suborder Anomaluromorpha

Family Anomaluridae

Members of the Anomaluridae family are called anomalurids or scaly-tailed squirrels. They eat a variety of vegetation, and some also eat insects.[4] Anomaluridae comprises seven extant species, divided into three genera.

Not assigned to a named subfamily – three genera
Name Authority and species Range Size and ecology
Anomalurus
(scaly-tailed flying squirrel)

Brown squirrel

Gervais, 1849

Four species
Sub-Saharan Africa (in red)
Map of range
Size range: 19 cm (7 in) long, plus 12 cm (5 in) tail (dwarf scaly-tailed squirrel) to 46 cm (18 in) long, plus 45 cm (18 in) tail (Pel's flying squirrel)[5]

Habitats: Forest[6]

Diets: Bark and fruit as well as flowers, leaves, nuts, and insects[4]
Idiurus
(flying mice)

Brown squirrel

Matschie, 1894

Two species
  • I. macrotis (Long-eared flying mouse, pictured)
  • I. zenkeri (Pygmy scaly-tailed flying squirrel)
Central and western Africa (in blue)
Map of range
Size range: 6 cm (2 in) long, plus 7 cm (3 in) tail (pygmy scaly-tailed flying squirrel) to 12 cm (5 in) long, plus 19 cm (7 in) tail (long-eared flying mouse)[5]

Habitats: Forest[7]

Diets: Fruit[4]
Zenkerella

Gray squirrel

Matschie, 1898

One species
West-central Africa (in purple)
Map of range
Size range: 18–23 cm (7–9 in) long, plus 15–18 cm (6–7 in) tail[5]

Habitats: Forest and savanna[8]

Diets: Vegetation[4]

Family Pedetidae

Members of the Pedetidae family are called pedetids or springhares. They eat bulbs and roots, as well as stems and grain.[9] Pedetidae comprises two extant species in a single genus.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Pedetes
(springhare)

Brown springhare

Illiger, 1811

Two species
Southern and eastern Africa
Map of range
Size range: 34–46 cm (13–18 in) long, plus 39–48 cm (15–19 in) tail[10]

Habitats: Savanna, shrubland, and grassland[11]

Diets: Bulbs and roots, as well as stems and grain[9]

Suborder Castorimorpha

Superfamily Castoroidea

Family Castoridae

Members of the Castoridae family are called castorids or beavers. They eat woody and aquatic plants.[12] Castoridae comprises two extant species in a single genus.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Castor
(beaver)

Brown springhare

Linnaeus, 1758

Two species
North America, Europe, and Asia
Map of range
Size range: 80–90 cm (31–35 in) long, plus 20–30 cm (8–12 in) tail[13]

Habitats: Forest, shrubland, and inland wetlands[14]

Diets: Bark, twigs, leaves, aquatic plants, and tree and shrub roots[12]

Superfamily Geomyoidea

Family Geomyidae

Members of the Geomyidae family are called geomyids, pocket gophers, or gophers. They primarily eat roots, bulbs, and rhizomes, as well as other vegetation.[15] Geomyidae comprises 36 extant species in 7 genera.

Not assigned to a named subfamily – seven genera
Name Authority and species Range Size and ecology
Cratogeomys
(pocket gopher)

Brown gopher

Merriam, 1895

Seven species
  • C. castanops (Yellow-faced pocket gopher, pictured)
  • C. fulvescens (Oriental Basin pocket gopher)
  • C. fumosus (Smoky pocket gopher)
  • C. goldmani (Goldman's pocket gopher)
  • C. merriami (Merriam's pocket gopher)
  • C. perotensis (Perote pocket gopher)
  • C. planiceps (Volcan de Toluca pocket gopher)
Southern United States and Mexico Size range: 14 cm (6 in) long, plus 5 cm (2 in) tail (Goldman's pocket gopher) to 28 cm (11 in) long, plus 13 cm (5 in) tail (Merriam's pocket gopher)[16]

Habitats: Grassland, shrubland, forest, and desert[17]

Diets: A variety of roots and vegetation, including cacti[15]
Geomys
(eastern pocket gopher)

Brown gopher

Rafinesque, 1817

Nine species
South-central Canada, United States, and Mexico Size range: 11 cm (4 in) long, plus 5 cm (2 in) tail (Knox Jones's pocket gopher) to 28 cm (11 in) long, plus 12 cm (5 in) tail (Texas pocket gopher)[16]

Habitats: Grassland, shrubland, savanna, forest, and desert[18]

Diets: Roots, rhizomes, and bulbs, as well as other vegetation[15]
Heterogeomys
(pocket gopher)
Le Conte, 1852

Three species
  • H. cherriei (Cherrie's pocket gopher)
  • H. dariensis (Darien pocket gopher)
  • H. lanius (Big pocket gopher)
Central America and Central Mexico Size range: 18 cm (7 in) long, plus 8 cm (3 in) tail (Cherrie's pocket gopher) to 29 cm (11 in) long, plus 10 cm (4 in) tail (big pocket gopher)[16]

Habitats: Forest[19]

Diets: A variety of vegetation[15]
Orthogeomys
(taltuza)

Brown gopher head

Merriam, 1895

Six species
  • O. cavator (Chiriqui pocket gopher)
  • O. cuniculus (Oaxacan pocket gopher)
  • O. grandis (Giant pocket gopher)
  • O. heterodus (Variable pocket gopher)
  • O. hispidus (Hispid pocket gopher, pictured)
  • O. underwoodi (Underwood's pocket gopher)
Mexico and Central America Size range: 18 cm (7 in) long, plus 8 cm (3 in) tail (Underwood's pocket gopher) to 30 cm (12 in) long, plus 14 cm (6 in) tail (giant pocket gopher)[16]

Habitats: Intertidal marine, forest, and grassland[20]

Diets: A variety of vegetation[15]
Pappogeomys Merriam, 1895

One species
Western-central Mexico Size: 12–18 cm (5–7 in) long, plus 6–10 cm (2–4 in) tail[16]

Habitats: Forest, shrubland, and grassland[21]

Diet: Roots and other vegetation[15]
Thomomys
(smooth-toothed pocket gopher)

Brown gopher

Wied-Neuwied, 1839

Nine species
Southwestern Canada, western United States, and northern Mexico Size range: 8 cm (3 in) long, plus 5 cm (2 in) tail (Botta's pocket gopher) to 24 cm (9 in) long, plus 10 cm (4 in) tail (Camas pocket gopher)[16]

Habitats: Inland wetlands, grassland, shrubland, savanna, forest, and desert[22]

Diets: Roots, bulbs, leaves, and other vegetation[15]
Zygogeomys Merriam, 1895

One species
Central Mexico Size: 17–24 cm (7–9 in) long, plus 6–10 cm (2–4 in) tail[16]

Habitats: Forest[23]

Diet: Roots and rhizomes[15]
Family Heteromyidae

Members of the Heteromyidae family are called heteromyids and include kangaroo rats, kangaroo mice, pocket mice, and spiny pocket mice. They primarily eat seeds, leaves, and other vegetation, as well as grain and insects.[24] Heteromyidae comprises 63 extant species in 5 genera. These genera are divided between three subfamilies: Dipodomyinae, containing the kangaroo rats and kangaroo mice; Heteromyinae, or the spiny pocket mice; and Perognathinae, or the pocket mice.

Subfamily Dipodomyinae – Gervais, 1853 – two genera
Name Authority and species Range Size and ecology
Dipodomys
(kangaroo rat)

Brown and white kangaroo rat

Gray, 1841

20 species
Southern Canada, western United States, and Mexico Size range: 7 cm (3 in) long, plus 13 cm (5 in) tail (Fresno kangaroo rat) to 16 cm (6 in) long, plus 21 cm (8 in) tail (desert kangaroo rat)[25]

Habitats: Rocky areas, shrubland, savanna, grassland, coastal marine, desert, and forest[26]

Diets: Seeds, as well as fruit, leaves, stems, buds, and insects[24]
Microdipodops
(kangaroo mouse)

Brown and white long-tailed rat

Merriam, 1891

Two species
  • M. megacephalus (Dark kangaroo mouse)
  • M. pallidus (Pale kangaroo mouse, pictured)
Western United States Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail to 7 cm (3 in) long, plus 11 cm (4 in) tail (dark kangaroo mouse)[25]

Habitats: Desert and shrubland[27]

Diets: Grains and insects[24]
Subfamily Heteromyinae – Gray, 1868 – one genus
Name Authority and species Range Size and ecology
Heteromys
(spiny pocket mouse)

Gray spiny pocket mouse

Desmarest, 1817

Fourteen species
  • H. adspersus (Panamanian spiny pocket mouse)
  • H. anomalus (Trinidad spiny pocket mouse)
  • H. australis (Southern spiny pocket mouse)
  • H. catopterius (Overlook spiny pocket mouse)
  • H. desmarestianus (Desmarest's spiny pocket mouse)
  • H. gaumeri (Gaumer's spiny pocket mouse)
  • H. irroratus (Mexican spiny pocket mouse)
  • H. nelsoni (Nelson's spiny pocket mouse, pictured)
  • H. oasicus (Paraguaná spiny pocket mouse)
  • H. oresterus (Mountain spiny pocket mouse)
  • H. pictus (Painted spiny pocket mouse)
  • H. salvini (Salvin's spiny pocket mouse)
  • H. spectabilis (Jaliscan spiny pocket mouse)
  • H. teleus (Ecuadoran spiny pocket mouse)
Southern United States, Mexico, Central America, and northeastern South America Size range: 10 cm (4 in) long, plus 12 cm (5 in) tail (Jaliscan spiny pocket mouse) to 17 cm (7 in) long, plus 20 cm (8 in) tail (Nelson's spiny pocket mouse)[25]

Habitats: Shrubland and forest[28]

Diets: Seeds, leaves, twigs, and succulent plants[24]
Subfamily PerognathinaeCoues, 1875 – two genera
Name Authority and species Range Size and ecology
Chaetodipus
(coarse-haired pocket mouse)

Gray pocket mouse

Merriam, 1889

Eighteen species
United States and Mexico Size range: 6 cm (2 in) long, plus 8 cm (3 in) tail (little desert pocket mouse) to 12 cm (5 in) long, plus 12 cm (5 in) tail (Hispid pocket mouse)[25]

Habitats: Rocky areas, shrubland, grassland, desert, and forest[29]

Diets: Seeds, as well as vegetation and insects[24]
Perognathus
(silky pocket mouse)

Brown pocket mouse

Wied-Neuwied, 1839

Nine species
Southern Canada, United States, and Mexico Size range: 5 cm (2 in) long, plus 4 cm (2 in) tail (Merriam's pocket mouse) to 8 cm (3 in) long, plus 9 cm (4 in) tail (white-eared pocket mouse)[25]

Habitats: Desert, shrubland, savanna, grassland, and forest[30]

Diets: Seeds, as well as vegetation and insects[24]

Suborder Hystricomorpha

Parvorder Caviomorpha

Superfamily Cavioidea
Family Caviidae

Members of the Caviidae family are called caviids and include guinea pigs, cavies, maras, and capybaras. They are herbivores and eat a wide variety of vegetation, primarily grass and leaves.[31] Caviidae comprises 21 extant species which are divided into 6 genera in 3 subfamilies: Caviinae, containing the guinea pigs and cavies; Dolichotinae, containing the maras; and Hydrochoerinae, containing rock cavies and capybaras.

Subfamily CaviinaeFischer de Waldheim, 1817 – three genera
Name Authority and species Range Size and ecology
Cavia
(guinea pig)

Black and white guinea pig

Pallas, 1766

Seven species
South America (guinea pig domesticated worldwide) Size range: 20 cm (8 in) long, with no tail (guinea pig) to 39 cm (15 in) long, with no tail (Brazilian guinea pig)[32]

Habitats: Grassland, forest, rocky areas, savanna, and inland wetlands[33]

Diets: Wide variety of vegetation[31]
Galea
(yellow-toothed cavy)

Brown cavy

Meyen, 1832

Five species
  • G. comes (Southern highland yellow-toothed cavy)
  • G. flavidens (Brazilian yellow-toothed cavy)
  • G. leucoblephara (Lowland yellow-toothed cavy)
  • G. musteloides (Common yellow-toothed cavy, pictured)
  • G. spixii (Spix's yellow-toothed cavy)
South America Size range: 19 cm (7 in) long, with no tail (lowland yellow-toothed cavy) to 24 cm (9 in) long, with no tail (southern highland yellow-toothed cavy)[32]

Habitats: Savanna, rocky areas, shrubland, and grassland[34]

Diets: Grass, forbs, and other vegetation[31]
Microcavia
(mountain cavy)

Brown cavy

Gervais & Ameghino, 1880

Three species
  • M. australis (Southern mountain cavy, pictured)
  • M. niata (Andean mountain cavy)
  • M. shiptoni (Shipton's mountain cavy)
Southern and western South America Size range: 19–20 cm (7–8 in) long, with no tail (Andean mountain cavy)[32]

Habitats: Desert, grassland, forest, rocky areas, savanna, and inland wetlands[35]

Diets: Leaves, as well as fruit[31]
Subfamily DolichotinaePocock, 1922 – one genus
Name Authority and species Range Size and ecology
Dolichotis
(mara)

Brown long-legged cavy

Desmarest, 1820

Two species
Southern and western South America Size range: 42 cm (17 in) long, plus 1 cm (0.4 in) tail (Chacoan mara) to 80 cm (31 in) long, plus 4 cm (2 in) tail (Patagonian mara)[32]

Habitats: Savanna, forest, shrubland, and grassland[36]

Diets: Wide variety of vegetation[31]
Subfamily Hydrochoerinae – J. E. Gray, 1825 – two genera
Name Authority and species Range Size and ecology
Hydrochoerus
(capybara)

Brown capybara

Brisson, 1762

Two species
Northwestern South America and Panama and South America
Map of range
Size range: 102 cm (40 in) long, plus 1 cm (0.4 in) tail (lesser capybara) to 134 cm (53 in) long, plus 2 cm (0.8 in) tail (capybara)[32]

Habitats: Forest, inland wetlands, and savanna[37]

Diets: Grass, as well as aquatic plants, grains, melons, and squashes[31]
Kerodon
(rock cavy)

Two brown cavies

F. Cuvier, 1825

Two species
  • K. acrobata (Acrobatic cavy)
  • K. rupestris (Rock cavy, pictured)
Brazil Size range: 30 cm (12 in) long, with no tail (rock cavy) to 38 cm (15 in) long, with no tail (acrobatic cavy)[32]

Habitats: Forest, rocky areas, and savanna[38]

Diets: Leaves, flowers, buds, bark, and occasionally fruit[31]
Family Cuniculidae

Members of the Cuniculidae family are called cuniculids or pacas. It contains two species, which eat leaves, stems, roots, seeds, and fruit.[39]

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Cuniculus
(paca)

Brown paca

Brisson, 1762

Two species
Mexico, Cuba, Central America, and South America
Map of range
Size range: 50 cm (20 in) long, plus 1 cm (0.4 in) tail (lowland paca) to 80 cm (31 in) long, plus 3 cm (1 in) tail (mountain paca)[40]

Habitats: Forest and inland wetlands[41]

Diets: Leaves, stems, roots, seeds, and fruit[39]
Family Dasyproctidae

Members of the Dasyproctidae family are called dasyproctids and include agoutis and acouchis. They are herbivores and primarily eat fruit, vegetables, and succulent plants.[42] Caviidae comprises fifteen extant species which are divided into two genera.

Not assigned to a named subfamily – two genera
Name Authority and species Range Size and ecology
Dasyprocta
(agouti)

Brown agouti

Bonaparte, 1838

Thirteen species
  • D. azarae (Azara's agouti)
  • D. coibae (Coiban agouti)
  • D. croconota (Orange agouti)
  • D. fuliginosa (Black agouti)
  • D. guamara (Orinoco agouti)
  • D. iacki (Iack's red-rumped agouti)
  • D. kalinowskii (Kalinowski's agouti)
  • D. leporina (Red-rumped agouti)
  • D. mexicana (Mexican agouti)
  • D. prymnolopha (Black-rumped agouti)
  • D. punctata (Central American agouti, pictured)
  • D. ruatanica (Ruatan Island agouti)
  • D. variegata (Brown agouti)
Mexico, Cuba, Central America, and South America (in red)
Map of range
Size range: 43 cm (17 in) long, plus 1 cm (0.4 in) tail (Azara's agouti) to 76 cm (30 in) long, plus 4 cm (2 in) tail (black agouti)[43]

Habitats: Savanna and forest[44]

Diets: Fruit, seeds, vegetables, and succulent plants, as well as crabs[42]
Myoprocta
(achouchi)

Brown achouchi

Thomas, 1903

Two species
  • M. acouchy (Red acouchi)
  • M. pratti (Green acouchi, pictured)
Northern South America (in blue)
Map of range
Size range: 30 cm (12 in) long, plus 4 cm (2 in) tail (green acouchi) to 39 cm (15 in) long, plus 8 cm (3 in) tail (red acouchi)[43]

Habitats: Forest[45]

Diets: Fruit, seeds, vegetables, and succulent plants[42]
Family Erethizontidae

Members of the Erethizontidae family are called erethizontids or New World porcupines. They are herbivores and primarily eat leaves, stems, fruit, flowers, and roots.[46] Erethizontidae comprises eighteen extant species in three genera, grouped into two subfamilies: Chaetomyinae, containing a single species, and Erethizontinae, containing the other seventeen.

Subfamily ChaetomyinaeThomas, 1897 – one genus
Name Authority and species Range Size and ecology
Chaetomys

Brown porcupine

J. E Gray, 1843

One species
Eastern Brazil
Map of range
Size: 36–45 cm (14–18 in) long, plus 26–27 cm (10–11 in) tail[47]

Habitats: Forest[48]

Diet: Nuts[46]
Subfamily ErethizontinaeBonaparte, 1845 – two genera
Name Authority and species Range Size and ecology
Coendou
(prehensile-tailed porcupine)

Black and yellow porcupine

Lacépède, 1799

Sixteen species
  • C. baturitensis (Baturite porcupine)
  • C. bicolor (Bicolored-spined porcupine)
  • C. ichillus (Streaked dwarf porcupine)
  • C. insidiosus (Bahia porcupine)
  • C. melanurus (Black-tailed hairy dwarf porcupine)
  • C. mexicanus (Mexican hairy dwarf porcupine)
  • C. nycthemera (Black dwarf porcupine)
  • C. prehensilis (Brazilian porcupine)
  • C. pruinosus (Frosted hairy dwarf porcupine)
  • C. quichua (Andean porcupine)
  • C. roosmalenorum (Roosmalen's dwarf porcupine)
  • C. rothschildi (Rothschild's porcupine)
  • C. rufescens (Stump-tailed porcupine)
  • C. speratus (Dwarf porcupine)
  • C. spinosus (Paraguaian hairy dwarf porcupine, pictured)
  • C. vestitus (Brown hairy dwarf porcupine)
Mexico, Central America, and South America
Map of range
Size range: 24 cm (9 in) long, plus 20 cm (8 in) tail (Paraguaian hairy dwarf porcupine) to 55 cm (22 in) long, plus 47 cm (19 in) tail (Baturite porcupine)[47]

Habitats: Forest and savanna[49]

Diets: Leaves, stems, fruit, flowers, and roots[46]
Erethizon

Brown and white porcupines

F. Cuvier, 1823

One species
North America
Map of range
Size: 60–130 cm (24–51 in) long, plus 16–25 cm (6–10 in) tail[47]

Habitats: Forest, shrubland, and grassland[50]

Diet: Buds, twigs, roots, stems, leaves, flowers, seeds, berries, nuts, and other vegetation[46]
Superfamily Chinchilloidea
Family Chinchillidae

Members of the Chinchillidae family are called chinchillids and contain chinchillas and viscachas. They are herbivores and eat grass and seeds, as well as other vegetation.[51] Chinchillidae comprises six extant species which are divided into three genera, as well as a single extinct species.

Not assigned to a named subfamily – three genera
Name Authority and species Range Size and ecology
Chinchilla
(chinchilla)

Gray chinchilla

Bennett, 1829

Two species
Western South America
Map of range
Size range: 22 cm (9 in) long, plus 14 cm (6 in) tail (long-tailed chinchilla) to 38 cm (15 in) long, plus 10 cm (4 in) tail (short-tailed chinchilla)[52]

Habitats: Grassland, shrubland, and rocky areas[53]

Diets: A wide variety of vegetation[51]
Lagidium
(viscacha)

Brown viscacha

Meyen, 1833

Three species
Western and southern South America
Map of range
Size range: 29 cm (11 in) long, plus 31 cm (12 in) tail (southern viscacha) to 48 cm (19 in) long, plus 31 cm (12 in) tail (Wolffsohn's viscacha)[52]

Habitats: Rocky areas[54]

Diets: A wide variety of vegetation, as well as lichen and moss[51]
Lagostomus
(plains viscacha)

Gray viscacha

Brookes, 1828

Two species
  • L. crassus (Peruvian plains viscacha)
  • L. maximus (Plains viscacha, pictured)
Southern South America
Map of range
Size range: 39–61 cm (15–24 in) long, plus 13–20 cm (5–8 in) tail (plains viscacha)[52]

Habitats: Savanna, grassland, and desert[55]

Diets: Grass and seeds, as well as other vegetation[51]
Family Dinomyidae

Members of the Dinomyidae family are called dinomyids. It contains a single species, which eats fruit, leaves, and stems.[56]

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Dinomys

Brown pacarana

Peters, 1873

One species
Northwestern South America
Map of range
Size: 73–79 cm (29–31 in) long, plus 14–23 cm (6–9 in) tail (pacarana)[57]

Habitats: Forest and grassland[58]

Diet: Fruit, leaves, and stems[56]
Superfamily Octodontoidea
Family Abrocomidae

Members of the Abrocomidae family are called abrocomids or chinchilla rats. They are herbivores and eat a variety of vegetation.[59] Abrocomidae comprises ten extant species which are divided into two genera.

Not assigned to a named subfamily – two genera
Name Authority and species Range Size and ecology
Abrocoma
(chinchilla rat)

Brown chinchilla rat

Waterhouse, 1837

Eight species
  • A. bennettii (Bennett's chinchilla rat, pictured)
  • A. boliviensis (Bolivian chinchilla rat)
  • A. budini (Budin's chinchilla rat)
  • A. cinerea (Ashy chinchilla rat)
  • A. famatina (Famatina chinchilla rat)
  • A. schistacea (Sierra del Tontal chinchilla rat)
  • A. uspallata (Uspallata chinchilla rat)
  • A. vaccarum (Punta de Vacas chinchilla rat)
Western South America (in red)
Map of range
Size range: 15 cm (6 in) long, plus 5 cm (2 in) tail (ashy chinchilla rat) to 23 cm (9 in) long, plus 28 cm (11 in) tail (Bennett's chinchilla rat)[57]

Habitats: Shrubland, inland wetlands, forest, rocky areas, grassland, and savanna[60]

Diets: A variety of vegetation[59]
Cuscomys
(arboreal chinchilla rat)

Drawing of chinchilla rat

Emmons, 1999

Two species
  • C. ashaninka (Asháninka arboreal chinchilla rat)
  • C. oblativus (Machu Picchu arboreal chinchilla rat, pictured)
Peru
Map of range
Size range: About 35 cm (14 in) long, plus about 26 cm (10 in) tail (Asháninka arboreal chinchilla rat)[57]

Habitats: Forest[61]

Diets: A variety of vegetation[59]
Family Capromyidae

Members of the Capromyidae family are called capromyids or hutias. They are omnivores and eat leaves, fruit, bark, and small animals.[62] Capromyidae comprises ten extant species in five genera, grouped into two subfamilies: Capromyinae, containing nine species, and Plagiodontinae, containing a single species. Additionally, two species in Capromyinae, three species in Plagiodontinae, the sole species in the subfamily Hexolobodontinae, and the two species in the subfamily Isolobodontinae were driven extinct after 1500 due to the European colonization of the Americas and introduction of non-native rats, with some species surviving until the 1900s.[63]

Subfamily CapromyinaeSmith, 1842 – four genera
Name Authority and species Range Size and ecology
Capromys

Brown hutia

Desmarest, 1822

One species
Cuba
Map of range
Size: 30–62 cm (12–24 in) long, plus 13–31 cm (5–12 in) tail[63]

Habitats: Forest and caves[64]

Diet: Leaves, fruit, bark, lizards, and other small animals[62]
Geocapromys
(Bahamian and Jamaican hutia)

Stuffed brown hutia

Chapman, 1901

Four species
  • G. brownii (Jamaican coney, pictured)
  • G. columbianus (Cuban coney)
  • G. ingrahami (Bahamian hutia)
  • G. thoracatus (Little Swan Island hutia)
Caribbean
Map of range
Size range: 28 cm (11 in) long, plus 6 cm (2 in) tail (Bahamian hutia) to 45 cm (18 in) long, plus 7 cm (3 in) tail (Jamaican coney)[63]

Habitats: Shrubland, intertidal marine, forest, and rocky areas[65]

Diets: Bark, twigs, and leaves[62]
Mesocapromys
(small Cuban hutia)

Brown hutia

Varona, 1970

Five species
  • M. angelcabrerai (Cabrera's hutia, pictured)
  • M. auritus (Eared hutia)
  • M. melanurus (Black-tailed hutia)
  • M. nanus (Dwarf hutia)
  • M. sanfelipensis (San Felipe hutia)
Cuba
Map of range
Size range: 20 cm (8 in) long, plus 17 cm (7 in) tail (Dwarf hutia) to 38 cm (15 in) long, plus 32 cm (13 in) tail (Black-tailed hutia)[63]

Habitats: Inland wetlands and forest[66]

Diets: Vegetation and small animals[62]
Mysateles

Stuffed brown hutia

Lesson, 1842

One species
Western Cuba
Map of range
Size: 33–45 cm (13–18 in) long, plus 25–35 cm (10–14 in) tail[63]

Habitats: Forest[67]

Diet: Leaves[67]
Subfamily Hexolobodontinae Woods, 1989 – one genus
Name Authority and species Range Size and ecology
Hexolobodon 

Hutia jawbone

Miller, 1929

One species
  • H. phenax (Imposter hutia)
Hispaniola Size: Unknown[63]

Habitats: Unknown[68]

Diet: Unknown[62]
Subfamily Isolobodontinae Woods, 1989 – one genus
Name Authority and species Range Size and ecology
Isolobodon 
(laminar-toothed hutia)

Hutia jawbone

J. A. Allen, 1916

Two species
  • I. montanus (Montane hutia)
  • I. portoricensis (Puerto Rican hutia, pictured)
Hispaniola, Puerto Rico, and nearby islands Size range: Unknown[63]

Habitats: Forest[69]

Diets: Unknown[62]
Subfamily PlagiodontinaeEllerman, 1940 – three genera
Name Authority and species Range Size and ecology
Hyperplagiodontia  Rímoli, 1977

One species
  • H. araeum (Wide-toothed hutia)
Hispaniola Size: Unknown[63]

Habitats: Unknown

Diet: Bark, leaves, and tree buds[62]
Plagiodontia
(Hispaniolan hutia)

Drawing of black hutia

F. Cuvier, 1836

Two species
Hispaniola
Map of range
Size range: 31–40 cm (12–16 in) long, plus 12–16 cm (5–6 in) tail (Hispaniolan hutia)[63]

Habitats: Rocky areas, and forest[70]

Diets: Bark, leaves, and tree buds[62]
Rhizoplagiodontia  Woods, 1989

One species
  • R. lemkei (Lemke's hutia)
Hispaniola Size: Unknown[63]

Habitats: Unknown

Diet: Unknown[62]
Family Ctenomyidae

Members of the Ctenomyidae family are called ctenomyids or tuco-tucos. They are herbivores and eat a wide variety of vegetation.[71] Ctenomyidae comprises 59 extant species in a single genus.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Ctenomys
(tuco-tuco)

Brown rodent

Blainville, 1826

59 species
  • C. argentinus (Argentine tuco-tuco)
  • C. australis (Southern tuco-tuco)
  • C. azarae (Azara's tuco-tuco)
  • C. bergi (Berg's tuco-tuco)
  • C. boliviensis (Bolivian tuco-tuco)
  • C. bonettoi (Bonetto's tuco-tuco)
  • C. brasiliensis (Brazilian tuco-tuco)
  • C. colburni (Colburn's tuco-tuco)
  • C. coludo (Puntilla tuco-tuco)
  • C. conoveri (Conover's tuco-tuco)
  • C. coyhaiquensis (Coyhaique tuco-tuco)
  • C. dorbignyi (D'Orbigny's tuco-tuco)
  • C. dorsalis (Chacoan tuco-tuco)
  • C. emilianus (Emily's tuco-tuco)
  • C. famosus (Famatina tuco-tuco)
  • C. flamarioni (Flamarion's tuco-tuco)
  • C. fochi (Foch's tuco-tuco)
  • C. fodax (Lago Blanco tuco-tuco)
  • C. frater (Reddish tuco-tuco)
  • C. fulvus (Tawny tuco-tuco)
  • C. goodfellowi (Goodfellow's tuco-tuco)
  • C. haigi (Haig's tuco-tuco)
  • C. ibicui (Ibicui tuco-tuco)
  • C. johannis (San Juan tuco-tuco)
  • C. juris (Jujuy tuco-tuco)
  • C. knighti (Catamarca tuco-tuco)
  • C. lami (Lami tuco-tuco)
  • C. latro (Mottled tuco-tuco)
  • C. leucodon (White-toothed tuco-tuco)
  • C. lewisi (Lewis's tuco-tuco)
  • C. magellanicus (Magellanic tuco-tuco)
  • C. maulinus (Maule tuco-tuco)
  • C. mendocinus (Mendoza tuco-tuco)
  • C. minutus (Tiny tuco-tuco)
  • C. occultus (Furtive tuco-tuco)
  • C. opimus (Highland tuco-tuco)
  • C. osvaldoreigi (Reig's tuco-tuco)
  • C. pearsoni (Pearson's tuco-tuco)
  • C. perrensi (Goya tuco-tuco)
  • C. peruanus (Peruvian tuco-tuco)
  • C. pilarensis (Pilar tuco-tuco)
  • C. pontifex (San Luis tuco-tuco)
  • C. porteousi (Porteous's tuco-tuco)
  • C. pundti (Pundt's tuco-tuco)
  • C. rionegrensis (Rio Negro tuco-tuco)
  • C. roigi (Roig's tuco-tuco)
  • C. saltarius (Salta tuco-tuco)
  • C. scagliai (Scaglia's tuco-tuco)
  • C. sericeus (Silky tuco-tuco, pictured)
  • C. sociabilis (Social tuco-tuco)
  • C. steinbachi (Steinbach's tuco-tuco)
  • C. talarum (Talas tuco-tuco)
  • C. torquatus (Collared tuco-tuco)
  • C. tuconax (Robust tuco-tuco)
  • C. tucumanus (Tucuman tuco-tuco)
  • C. tulduco (Sierra Tontal tuco-tuco)
  • C. validus (Strong tuco-tuco)
  • C. viperinus (Vipos tuco-tuco)
  • C. yolandae (Yolanda's tuco-tuco)
Southern South America
Map of range
Size range: 11 cm (4 in) long, plus 7 cm (3 in) tail (white-toothed tuco-tuco) to 33 cm (13 in) long, plus 14 cm (6 in) tail (Conover's tuco-tuco)[72]

Habitats: Forest, desert, grassland, inland wetlands, coastal marine, shrubland, and savanna[73]

Diets: A wide variety of vegetation[71]
Family Echimyidae

Members of the Echimyidae family are called echimyids or Neotropical spiny rats, and include spiny rats, tree-rats, and cave rats. They primarily eat a wide variety of vegetation, though some species also eat insects.[74] Echimyidae comprises 88 extant species in 23 genera, split into three subfamilies: Caterodontinae, containing a single species; Echimyinae, containing 75 species, and Euryzygomatomyinae, containing 12 species. Additionally, six species of Caribbean cave rat, comprising the three genera of the subfamily Heteropsomyinae, were driven to extinction since 1500 due to the European colonization of the Americas and introduction of non-native rats, with some species surviving until the 1900s.[75]

Subfamily Carterodontinae – Courcelle, Tilak, Leite, Douzery, & Fabre, 2019 – one genus
Name Authority and species Range Size and ecology
Carterodon Waterhouse, 1848

One species
  • C. sulcidens (Owl's spiny rat)
Eastern Brazil Size: 13–25 cm (5–10 in) long, plus 6–10 cm (2–4 in) tail[75]

Habitats: Forest and savanna[76]

Diet: Vegetation[74]
Subfamily Echimyinae – Gray, 1825 – eighteen genera
Name Authority and species Range Size and ecology
Callistomys

Gray and white spiny rat

Emmons & Vucetich, 1998

One species
Eastern Brazil Size: 25–29 cm (10–11 in) long, plus 27–32 cm (11–13 in) tail[75]

Habitats: Forest[77]

Diet: Vegetation[74]
Dactylomys
(bamboo rat)

Drawing of gray spiny rat

Geoffroy, 1838

Three species
Northern and western South America Size range: 24 cm (9 in) long, plus 32 cm (13 in) tail (montane bamboo rat) to 31 cm (12 in) long, plus 39 cm (15 in) tail (Amazon bamboo rat)[75]

Habitats: Forest[78]

Diets: Bamboo and other plants[74]
Diplomys
(soft-furred spiny rat)

Drawing of brown spiny rat

Thomas, 1916

Two species
  • D. caniceps (Colombian soft-furred spiny rat, pictured)
  • D. labilis (Rufous soft-furred spiny rat)
Northwestern South America and Panama Size range: 21–39 cm (8–15 in) long, plus 17–27 cm (7–11 in) tail (Colombian soft-furred spiny rat)[75]

Habitats: Forest[79]

Diets: Seeds and fruit[74]
Echimys
(spiny tree-rat)

Brown and white spiny rat

F. Cuvier, 1809

Three species
  • E. chrysurus (White-faced spiny tree-rat, pictured)
  • E. saturnus (Dark spiny tree-rat)
  • E. vieirai (Vieira's tree-rat)
Northern South America Size range: 24 cm (9 in) long, plus 34 cm (13 in) tail (Vieira's tree-rat) to 33 cm (13 in) long, plus 38 cm (15 in) tail (dark spiny tree-rat)[75]

Habitats: Forest[80]

Diets: Vegetation[74]
Hoplomys

Brown spiny rat

J. A. Allen, 1908

One species
Central America and northwestern South America Size: 21–32 cm (8–13 in) long, plus 11–24 cm (4–9 in) tail[75]

Habitats: Forest[81]

Diet: Wide variety of vegetation, as well as insects[74]
Isothrix
(brush-tailed rat)

Brown spiny rat

Wagner, 1845

Six species
  • I. barbarabrownae (Barbara Brown's brush-tailed rat)
  • I. bistriata (Yellow-crowned brush-tailed rat, pictured)
  • I. negrensis (Rio Negro brush-tailed rat)
  • I. orinoci (Orinoco brush-tailed rat)
  • I. pagurus (Plain brush-tailed rat)
  • I. sinnamariensis (Sinnamary brush-tailed rat)
Northern and western South America Size range: 17 cm (7 in) long, plus 17 cm (7 in) tail (Plain brush-tailed rat) to 29 cm (11 in) long, plus 27 cm (11 in) tail (Rio Negro brush-tailed rat)[75]

Habitats: Forest and grassland[82]

Diets: Vegetation[74]
Kannabateomys

Brown and white spiny rat

Jentink, 1891

One species
Eastern South America Size: 23–35 cm (9–14 in) long, plus 30–42 cm (12–17 in) tail[75]

Habitats: Forest[83]

Diet: Bamboo[74]
Lonchothrix Thomas, 1920

One species
Northern Brazil Size: 15–22 cm (6–9 in) long, plus 15–23 cm (6–9 in) tail[75]

Habitats: Forest and grassland[84]

Diet: Vegetation[74]
Makalata
(armored tree-rat)

Brown spiny rat

Husson, 1978

Three species
  • M. didelphoides (Brazilian spiny tree-rat, pictured)
  • M. macrura (Long-tailed armored tree-rat)
  • M. obscura (Dusky spiny tree-rat)
Northern South America Size range: 15 cm (6 in) long, plus 15 cm (6 in) tail (Brazilian spiny tree-rat) to 28 cm (11 in) long, plus 24 cm (9 in) tail (long-tailed armored tree-rat)[75]

Habitats: Forest[85]

Diets: Fruit and seeds[74]
Mesomys
(spiny tree-rat)

Brown spiny rat

Wagner, 1845

Four species
  • M. hispidus (Ferreira's spiny tree-rat, pictured)
  • M. leniceps (Woolly-headed spiny tree-rat)
  • M. occultus (Tufted-tailed spiny tree-rat)
  • M. stimulax (Pará spiny tree-rat)
Northern South America Size range: 15–20 cm (6–8 in) long, plus 12–20 cm (5–8 in) tail (Pará spiny tree-rat)[75]

Habitats: Forest[86]

Diets: Vegetation[74]
Myocastor

Brown nutria in water

Kerr, 1792

One species
Southern South America, and introduced in United States, Europe, and Japan
Map of range
Size: 47–57 cm (19–22 in) long, plus 34–40 cm (13–16 in) tail[75]

Habitats: Inland wetlands[87]

Diet: Roots and aquatic plants[74]
Olallamys
(olalla rat)

Drawing of brown and white spiny rat

Emmons, 1988

Two species
  • O. albicauda (White-tailed olalla rat, pictured)
  • O. edax (Greedy olalla rat)
Colombia and western Venezuela Size range: 15 cm (6 in) long, plus 25 cm (10 in) tail (white-tailed olalla rat) to 22 cm (9 in) long, plus 34 cm (13 in) tail (greedy olalla rat)[75]

Habitats: Forest and shrubland[88]

Diets: Vegetation[74]
Pattonomys
(speckled tree-rat)
Emmons, 2005

Five species
  • P. carrikeri (Carriker's speckled tree-rat)
  • P. flavidus (Yellow speckled tree-rat)
  • P. occasius (Bare-tailed armored tree-rat)
  • P. punctatus (Orinocoan speckled tree-rat)
  • P. semivillosus (Speckled spiny tree-rat)
Colombia, Venezuela, and Ecuador Size range: 20 cm (8 in) long, plus 21 cm (8 in) tail (speckled spiny tree-rat) to 27 cm (11 in) long, plus 23 cm (9 in) tail (Orinocoan speckled tree-rat)[75]

Habitats: Forest and savanna[89]

Diets: Vegetation[74]
Phyllomys
(Atlantic tree-rat)

Brown spiny rat

Lund, 1839

Thirteen species
  • P. blainvillii (Golden Atlantic tree-rat)
  • P. brasiliensis (Orange-brown Atlantic tree-rat)
  • P. dasythrix (Drab Atlantic tree-rat)
  • P. kerri (Kerr's Atlantic tree-rat)
  • P. lamarum (Pallid Atlantic tree-rat)
  • P. lundi (Lund's Atlantic tree-rat)
  • P. mantiqueirensis (Mantiqueira Atlantic tree-rat)
  • P. medius (Long-furred Atlantic tree-rat)
  • P. nigrispinus (Black-spined Atlantic tree-rat)
  • P. pattoni (Rusty-sided Atlantic tree-rat)
  • P. sulinus (Southern Atlantic tree-rat)
  • P. thomasi (Giant Atlantic tree-rat, pictured)
  • P. unicolor (Short-furred Atlantic tree-rat)
Southern and eastern Brazil Size range: 17 cm (7 in) long, plus 19 cm (7 in) tail (golden Atlantic tree-rat) to 30 cm (12 in) long, plus 34 cm (13 in) tail (giant Atlantic tree-rat)[75]

Habitats: Forest[90]

Diets: Vegetation[74]
Proechimys
(spiny rat)

Brown spiny rat

Allen, 1899

22 species
  • P. brevicauda (Short-tailed spiny rat)
  • P. canicollis (Colombian spiny rat)
  • P. chrysaeolus (Boyacá spiny rat)
  • P. cuvieri (Cuvier's spiny rat)
  • P. decumanus (Pacific spiny rat)
  • P. echinothrix (Stiff-spine spiny rat)
  • P. gardneri (Gardner's spiny rat)
  • P. goeldii (Goeldi's spiny rat)
  • P. guairae (Guaira spiny rat)
  • P. guyannensis (Guyenne spiny rat)
  • P. hoplomyoides (Guyanan spiny-rat)
  • P. kulinae (Kulina spiny rat)
  • P. longicaudatus (Long-tailed spiny rat, pictured)
  • P. mincae (Minca spiny rat)
  • P. oconnelli (O'Connell's spiny rat)
  • P. pattoni (Patton's spiny rat)
  • P. quadruplicatus (Napo spiny rat)
  • P. roberti (Robert's spiny rat)
  • P. semispinosus (Tome's spiny rat)
  • P. simonsi (Simons's spiny rat)
  • P. steerei (Steere's spiny rat)
  • P. trinitatus (Trinidad spiny rat)
Central America and Northern and western South America Size range: 14 cm (6 in) long, plus 9 cm (4 in) tail (Kulina spiny rat) to 30 cm (12 in) long, plus 20 cm (8 in) tail (Pacific spiny rat)[75]

Habitats: Forest and savanna[91]

Diets: Wide variety of vegetation[74]
Santamartamys

Red, brown, and white spiny rat

Emmons, 2005

One species
Northern Colombia
Map of range
Size: About 19 cm (7 in)[75]

Habitats: Unknown[92]

Diet: Seeds and fruit[74]
Thrichomys
(punaré)

Gray spiny rat

Trouessart, 1880

Four species
  • T. apereoides (Common punaré, pictured)
  • T. inermis (Highlands punaré)
  • T. laurentius (Sao Lourenço punaré)
  • T. pachyurus (Paraguayan punaré)
Southern and eastern Brazil Size range: 12 cm (5 in) long, plus 13 cm (5 in) tail (Sao Lourenço punaré) to 24 cm (9 in) long, plus 21 cm (8 in) tail (Sao Lourenço punaré)[75]

Habitats: Forest, rocky areas, savanna, and shrubland[93]

Diets: Seeds, fruit, cacti, and insects[74]
Toromys
(giant tree-rat)
Iack-Ximenes, Vivo, & Percequillo, 2005

Two species
Northern Brazil and Peru Size range: 21 cm (8 in) long, plus 18 cm (7 in) tail (Peruvian toro) to 30 cm (12 in) long, plus 28 cm (11 in) tail (giant tree-rat)[75]

Habitats: Forest[94]

Diets: Vegetation[74]
Subfamily Euryzygomatomyinae – Fabre, Upham, Emmons, Justy, Leite, Loss, Orlando, Tilak, Patterson, & Douzery, 2017 – three genera
Name Authority and species Range Size and ecology
Clyomys Thomas, 1916

One species
Paraguay and southern Brazil Size: 15–29 cm (6–11 in) long, plus 4–9 cm (2–4 in) tail[75]

Habitats: Savanna[95]

Diet: Vegetation[74]
Euryzygomatomys

Drawing of brown spiny rat

Goeldi, 1901

One species
  • E. spinosus (Fischer's guiara)
Southern Brazil and northeastern Argentina Size: 16–20 cm (6–8 in) long, plus 6–7 cm (2–3 in) tail[75]

Habitats: Forest and grassland[96]

Diet: Vegetation[74]
Trinomys
(Atlantic spiny rat)

Brown and white spiny rat

Thomas, 1921

Ten species
  • T. albispinus (White-spined Atlantic spiny rat)
  • T. dimidiatus (Soft-spined Atlantic spiny rat)
  • T. eliasi (Elias's Atlantic spiny rat)
  • T. gratiosus (Gracile Atlantic spiny rat)
  • T. iheringi (Ihering's Atlantic spiny rat)
  • T. mirapitanga (Dark-caped Atlantic spiny rat)
  • T. moojeni (Moojen's Atlantic spiny rat)
  • T. paratus (Spiked Atlantic spiny rat)
  • T. setosus (Hairy Atlantic spiny rat)
  • T. yonenagae (Yonenaga's Atlantic spiny rat)
Eastern Brazil Size range: 14 cm (6 in) long, plus 16 cm (6 in) tail (Yonenaga's Atlantic spiny rat) to 27 cm (11 in) long, plus 23 cm (9 in) tail (spiked Atlantic spiny rat)[75]

Habitats: Forest, savanna, rocky areas, grassland, and shrubland[97]

Diets: Wide variety of vegetation[74]
Subfamily HeteropsomyinaeAnthony, 1917 – three genera
Name Authority and species Range Size and ecology
Boromys 
(Cuban cave rat)
Miller, 1916

Two species
  • B. offella (Oriente cave rat)
  • B. torrei (Torre's cave rat)
Cuba Size range: Unknown[75]

Habitats: Unknown[98]

Diets: Unknown[74]
Brotomys 
(edible rat)
Miller, 1916

Two species
  • B. contractus (Haitian edible rat)
  • B. voratus (Hispaniolan edible rat)
Dominican Republic and Haiti Size range: Unknown[75]

Habitats: Unknown[99]

Diets: Unknown[74]
Heteropsomys 
(Puerto Rican cave rat)
Anthony, 1916

Two species
  • H. antillensis (Antillean cave rat)
  • H. insulans (Insular cave rat)
Puerto Rico Size range: Unknown[75]

Habitats: Unknown[100]

Diets: Unknown[74]
Family Octodontidae

Members of the Octodontidae family are called octodontids and include degus, rock rats, and viscacha rats. They are herbivores, and eat a variety of plant matter and fruit, with some species also eating dung.[101] Octodontidae comprises fourteen extant species in seven genera.

Not assigned to a named subfamily – seven genera
Name Authority and species Range Size and ecology
Aconaemys
(rock rat)

Illustration of a rock rat

Ameghino, 1891

Three species
  • A. fuscus (Chilean rock rat, pictured)
  • A. porteri (Porter's rock rat)
  • A. sagei (Sage's rock rat)
Chile and western Argentina Size range: 14–20 cm (6–8 in) long, plus 6–9 cm (2–4 in) tail (Porter's rock rat)[102]

Habitats: Rocky areas and forest[103]

Diets: Roots, fruit, and other vegetation[101]
Octodon
(degu)

A degu with a paw raised, viewed head-on

Bennett, 1823

Four species
  • O. bridgesii (Bridges's degu)
  • O. degus (Common degu, pictured)
  • O. lunatus (Moon-toothed degu)
  • O. pacificus (Pacific degu)
Chile Size range: 12 cm (5 in) long, plus 16 cm (6 in) tail (Pacific degu) to 22 cm (9 in) long, plus 17 cm (7 in) tail (moon-toothed degu)[102]

Habitats: Rocky areas, forest, and shrubland[104]

Diets: Grass, leaves, bark, herbs, seeds, and fruit, as well as dung[101]
Octodontomys

A degu obscured by sticks and grass

Palmer, 1903

One species
Western South America
Map of range
Size: 16–19 cm (6–7 in) long, plus 10–19 cm (4–7 in) tail[102]

Habitats: Shrubland, grassland, and rocky areas[105]

Diet: Acacia seed pods and cactus fruits[101]
Octomys Thomas, 1920

One species
Western Argentina Size: 14–33 cm (6–13 in) long, plus 10–18 cm (4–7 in) tail[102]

Habitats: Rocky areas[106]

Diet: Roots, bark, and cactus[101]
Pipanacoctomys Mares, Braun, Barquez, & Díaz, 2000

One species
Northwestern Argentina Size: 16–18 cm (6–7 in) long, plus 12–15 cm (5–6 in) tail[102]

Habitats: Inland wetlands[107]

Diet: Leaves and stems[101]
Spalacopus

Wagler, 1832

One species
Central Chile Size: 11–17 cm (4–7 in) long, plus 4–6 cm (2 in) tail[102]

Habitats: Forest, savanna and grassland[108]

Diet: Tubers and stems of Leucocoryne and other plants[101]
Tympanoctomys
(viscacha rat)

A rat with a hairy tail against a smooth white backdrop

Yepes, 1942

Three species
  • T. barrerae (Plains viscacha rat, pictured)
  • T. kirchnerorum (Kirchner's viscacha rat)
  • T. loschalchalerosorum (Chalchalero viscacha rat)
Argentina Size range: 11 cm (4 in) long, plus 11 cm (4 in) tail (Kirchner's viscacha rat) to 16 cm (6 in) long, plus 12 cm (5 in) tail (Chalchalero viscacha rat)[102]

Habitats: Grassland, shrubland, shrublands, and inland wetlands[109]

Diets: Leaves and stems[101]

Superfamily Ctenodactyloidea

Family Ctenodactylidae

Members of the Ctenodactylidae family are called ctenodactylids or gundis. They eat a variety of vegetation.[110] Ctenodactylidae comprises five extant species in four genera.

Not assigned to a named subfamily – four genera
Name Authority and species Range Size and ecology
Ctenodactylus
(gundi)

Brown gundi

J. E. Gray, 1830

Two species
Northwestern Africa Size range: 12 cm (5 in) long, plus 2 cm (0.8 in) tail (Val's gundi) to 23 cm (9 in) long, plus 5 cm (2 in) tail (common gundi)[111]

Habitats: Shrubland and rocky areas[112]

Diets: Vegetation[110]
Felovia

Brown gundi

Lastaste, 1886

One species
  • F. vae (Felou gundi)
Western Africa
Map of range
Size: 16–19 cm (6–7 in) long, plus 6–8 cm (2–3 in) tail[111]

Habitats: Savanna, shrubland, and rocky areas[113]

Diet: Vegetation[110]
Massoutiera

Drawing of brown gundi

Lastaste, 1885

One species
  • M. mzabi (Mzab gundi)
Northwestern Africa
Map of range
Size: 12–21 cm (5–8 in) long, plus 3–9 cm (1–4 in) tail[111]

Habitats: Rocky areas[114]

Diet: Leaves, stems, seeds, and flowers[110]
Pectinator

Drawing of brown gundi

Blyth, 1856

One species
  • P. spekei (Speke's pectinator)
Eastern Africa
Map of range
Size: 15–19 cm (6–7 in) long, plus 6–8 cm (2–3 in) tail[111]

Habitats: Shrubland, grassland, and rocky areas[115]

Diet: Leaves, stems, seeds, and flowers[110]
Family Diatomyidae

Members of the Diatomyidae family are called diatomyids. The family contains a single herbivorous species.[116]

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Laonastes

Gray rock rat

Jenkins, Kilpatrick, Robinson, & Timmins, 2005

One species
Laos and Vietnam
Map of range
Size: 21–30 cm (8–12 in) long, plus 12–17 cm (5–7 in) tail[111]

Habitats: Forest and rocky areas[116]

Diet: Grass, seeds, and other vegetation[116]

Family Hystricidae

Members of the Hystricidae family are called hystricids or Old World porcupines. They are primarily herbivores and eat grasses, bulbs, tubers, fruit, bark, and roots, though insects and carrion are occasionally eaten.[117] Hystricidae comprises eleven extant species in three genera.

Not assigned to a named subfamily – three genera
Name Authority and species Range Size and ecology
Atherurus
(brush-tailed porcupine)

Brown porcupine

F. Cuvier, 1829

Two species
Southeast Asia, southern China and Central Africa (outlined in blue)
Map of range
Size range: 36–60 cm (14–24 in) long, plus 10–26 cm (4–10 in) tail (African brush-tailed porcupine)[118]

Habitats: Forest and shrubland[119]

Diets: Vegetation, bark, roots, tubers, and fruit, as well as cultivated crops, insects, and carrion[117]
Hystrix
(Old World porcupine)

Brown and white porcupine

Linnaeus, 1758

Eight species
Africa, Italy, and Asia (solid colors)
Map of range
Size range: 45–93 cm (18–37 in) long, plus 6–17 cm (2–7 in) tail (crested porcupine)[118]

Habitats: Forest, grassland, shrubland, rocky areas, and savanna[120]

Diets: Bark, roots, tubers, rhizomes, bulbs, fruit, and cultivated crops, as well as carrion, insects, and small vertebrates[117]
Trichys

Brown porcupine under a desk

Günther, 1877

One species
Brunei, Indonesia, and Malaysia
Map of range
Size: 35–48 cm (14–19 in) long, plus 17–24 cm (7–9 in) tail[118]

Habitats: Forest[121]

Diet: Seeds, vegetation from trees, shrubs, and bamboo[118]

Parvorder Phiomorpha

Family Bathyergidae

Members of the Bathyergidae family are called bathyergids, blesmols, or mole-rats. They are omnivores, and primarily eat roots, bulbs, and tubers, as well as earthworms and insects.[122] Bathyergidae comprises twenty-one extant species in five genera.

Not assigned to a named subfamily – five genera
Name Authority and species Range Size and ecology
Bathyergus
(dune mole-rat)

Brown mole-rat

Illiger, 1811

Two species
South Africa and Namibia Size range: 17 cm (7 in) long, plus 4 cm (2 in) tail (Namaqua dune mole-rat) to 35 cm (14 in) long, plus 4 cm (2 in) tail (Cape dune mole-rat)[123]

Habitats: Desert and shrubland[124]

Diets: Bulbs and fleshy roots[122]
Cryptomys
(common mole-rat)

Brown mole-rat

Gray, 1864

Four species
  • C. hottentotus (Common mole-rat, pictured)
  • C. mahali (Mahali mole-rat)
  • C. natalensis (Natal mole-rat)
  • C. pretoriae (Highveld mole-rat)
South Africa, Lesotho, and Eswatini Size range: 13 cm (5 in) long, plus 1 cm (0.4 in) tail (Mahali mole-rat) to 18 cm (7 in) long, plus 3 cm (1 in) tail (Natal mole-rat)[123]

Habitats: Grassland and shrubland[125]

Diets: Roots, bulbs, tubers, and aloe leaves, as well as earthworms and insects[122]
Fukomys
(tropical mole-rat)

Brown mole-rat

Kock, Ingram, Frabotta, Honeycutt, & Burda, 2006

Thirteen species
  • F. anselli (Ansell's mole-rat)
  • F. bocagei (Bocage's mole-rat)
  • F. damarensis (Damaraland mole-rat, pictured)
  • F. darlingi (Mashona mole-rat)
  • F. foxi (Nigerian mole-rat)
  • F. hanangensis (Hanang mole-rat)
  • F. ilariae (Somali striped mole-rat)
  • F. kafuensis (Kafue mole-rat)
  • F. livingstoni (Livingstone's mole-rat)
  • F. mechowi (Mechow's mole-rat)
  • F. ochraceocinereus (Ochre mole-rat)
  • F. vandewoestijneae (Caroline's mole-rat)
  • F. zechi (Ghana mole-rat)
Central and southern Africa Size range: 8 cm (3 in) long, plus 1 cm (0.4 in) tail (Caroline's mole-rat) to 26 cm (10 in) long, plus 4 cm (2 in) tail (Mechow's mole-rat)[123]

Habitats: Grassland, savanna, forest, and shrubland[126]

Diets: Roots, bulbs, tubers, and aloe leaves, as well as earthworms and insects[122]
Georychus

Brown mole-rat

Illiger, 1811

One species
South Africa and Lesotho Size: 8–23 cm (3–9 in) long, plus 1–3 cm (0.4–1.2 in) tail[123]

Habitats: Shrubland and forest[127]

Diet: Tubers, roots, and bulbs[122]
Heliophobius

Brown mole-rat

Peters, 1846

One species
Southeastern Africa Size: 13–20 cm (5–8 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[123]

Habitats: Savanna, shrubland, grassland, and caves[128]

Diet: Tubers and bulbs[122]
Family Heterocephalidae

Members of the Heterocephalidae family are called heterocephalids. It includes a single species, which eats tubers, roots, and corms.[129]

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Heterocephalus

White mole rat

Rüppell, 1842

One species
Eastern Africa
Map of range
Size: 7–11 cm (3–4 in) long, plus 3–5 cm (1–2 in) tail[130]

Habitats: Savanna, shrubland, grassland, and caves[131]

Diet: Tubers, roots, and corms[129]
Family Petromuridae

Members of the Petromuridae family are called petromurids. It contains a single species, which eats vegetation, seeds, and berries.[132]

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Petromus

Gray dassie rat

A. Smith, 1831

One species
Southwestern Africa
Map of range
Size: 13–22 cm (5–9 in) long, plus 11–18 cm (4–7 in) tail[133]

Habitats: Shrubland and rocky areas[134]

Diet: Vegetation, seeds, and berries[132]
Family Thryonomyidae

Members of the Thryonomyidae family are called thryonomyids or cane rats. It contains two species, which eats grass and cane, as well as bark, nuts, and fruit.[135]

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Thryonomys
(cane rat)

Brown cane rat

Fitzinger, 1867

Two species
Sub-Saharan Africa
Map of range
Size range: 41 cm (16 in) long, plus 11 cm (4 in) tail (lesser cane rat) to 77 cm (30 in) long, plus 20 cm (8 in) tail (greater cane rat)[133]

Habitats: Shrubland, grassland, inland wetlands, and forest[136]

Diets: Grass and cane, as well as bark, nuts, and fruit[135]

Suborder Myomorpha

Superfamily Dipodoidea

Family Dipodidae

Members of the Dipodidae family are called dipodids or jerboas. They are omnivores, and eat a variety of vegetation as well as insects, arachnids, and lizards.[137] Dipodidae comprises 33 extant species in 13 genera, which are divided into 4 subfamilies: Allactaginae contains 16 species, Cardiocraniinae contains 7 species, Dipodinae contains 9, and Euchoreutinae contains a single species.

Subfamily Allactaginae – Vinogradov, 1925 – four genera
Name Authority and species Range Size and ecology
Allactaga
(five-toed jerboa)

Brown jerboa

F. Cuvier, 1836

Eight species
Northern Africa, central and western Asia, and eastern Europe Size range: 9 cm (4 in) long, plus 16 cm (6 in) tail (Hotson's jerboa) to 23 cm (9 in) long, plus 30 cm (12 in) tail (great jerboa)[138]

Habitats: Desert, coastal marine, shrubland, grassland, and forest[139]

Diets: Vegetation, seeds, roots, tubers, insects, and larvae, as well as molluscs[139]
Allactodipus Kolesnikov, 1937

One species
Central Asia Size: 11–14 cm (4–6 in) long, plus 17–20 cm (7–8 in) tail[138]

Habitats: Desert[140]

Diet: Vegetation and insects[140]
Pygeretmus
(fat-tailed jerboa)
Gloger, 1841

Three species
  • P. platyurus (Lesser fat-tailed jerboa)
  • P. pumilio (Dwarf fat-tailed jerboa)
  • P. shitkovi (Greater fat-tailed jerboa)
Central Asia Size range: 7 cm (3 in) long, plus 7 cm (3 in) tail (lesser fat-tailed jerboa) to 14 cm (6 in) long, plus 18 cm (7 in) tail (dwarf fat-tailed jerboa)[138]

Habitats: Shrubland and desert[141]

Diets: Succulent plants and other vegetation, bulbs, rhizomes, roots, spiders, and insects[137]
Scarturus
(four- and five-toed jerboa)

Brown jerboa

Gloger, 1841

Four species
Central and western Asia Size range: 7 cm (3 in) long, plus 14 cm (6 in) tail (Euphrates jerboa) to 15 cm (6 in) long, plus 26 cm (10 in) tail (Williams's jerboa)[138]

Habitats: Shrubland, grassland, and desert[142]

Diets: Sprouts, seeds, underground plant parts, other vegetation, and insects[137]
Subfamily Cardiocraniinae – Vinogradov, 1925 – three genera
Name Authority and species Range Size and ecology
Cardiocranius

Brown jerboa

Satunin, 1903

One species
East-central Asia Size: 5–7 cm (2–3 in) long, plus 6–10 cm (2–4 in) tail[138]

Habitats: Desert[143]

Diet: Seeds[137]
Salpingotulus

Brown jerboas

Pavlinov, 1980

One species
Pakistan Size: 4–5 cm (2 in) long, plus 7–10 cm (3–4 in) tail[138]

Habitats: Desert[144]

Diet: Grass seeds, stems, and other vegetation[137]
Salpingotus
(three-toed dwarf jerboa)

Brown jerboa

Vinogradov, 1922

Five species
Central Asia Size range: 4 cm (2 in) long, plus 8 cm (3 in) tail (Heptner's pygmy jerboa) to 7 cm (3 in) long, plus 14 cm (6 in) tail (Kozlov's pygmy jerboa)[138]

Habitats: Shrubland, desert, and grassland[145]

Diets: Insects, arachnids, and vegetation[137]
Subfamily Dipodinae – G. Fischer, 1817 – five genera
Name Authority and species Range Size and ecology
Dipus

Brown jerboa

Zimmermann, 1780

One species
Western, central, and eastern Asia Size: 10–15 cm (4–6 in) long, plus 12–20 cm (5–8 in) tail[138]

Habitats: Shrubland and desert[146]

Diet: All parts of plants, as well as insects[137]
Eremodipus Vinogradov, 1930

One species
Central Asia Size: 10–12 cm (4–5 in) long, plus 13–17 cm (5–7 in) tail[138]

Habitats: Desert[147]

Diet: Roots, sprouts, seeds, grains, and vegetables[137]
Jaculus
(desert jerboa)

Brown jerboa

Erxleben, 1777

Three species
Northern Africa, Middle East, and central Asia Size range: 9 cm (4 in) long, plus 17 cm (7 in) tail (greater Egyptian jerboa) to 15 cm (6 in) long, plus 22 cm (9 in) tail (Blanford's jerboa)[138]

Habitats: Desert, coastal marine, shrubland, grassland, and rocky areas[148]

Diets: Roots, sprouts, seeds, grains, and vegetables[137]
Paradipus Vinogradov, 1930

One species
Central Asia Size: 14–17 cm (6–7 in) long, plus 18–22 cm (7–9 in) tail[138]

Habitats: Desert[149]

Diet: All parts of desert plants[137]
Stylodipus
(thick-tailed three-toed jerboa)

Brown jerboa

Allen, 1925

Three species
Ukraine and western and central Asia Size range: 11 cm (4 in) long, plus 13 cm (5 in) tail (thick-tailed three-toed jerboa) to 14 cm (6 in) long, plus 17 cm (7 in) tail (Andrews's three-toed jerboa)[138]

Habitats: Shrubland, desert, grassland, and forest[150]

Diets: Lichen, rhizomes, bulbs, seeds, and wheat[137]
Subfamily EuchoreutinaeLyon, 1901 – one genus
Name Authority and species Range Size and ecology
Euchoreutes

Drawing of brown jerboa

W. L. Sclater, 1891

One species
Southern Mongolia and China Size: 9–11 cm (4 in) long, plus 14–18 cm (6–7 in) tail[138]

Habitats: Shrubland and desert[151]

Diet: Insects and lizards[151]
Family Sminthidae

Members of the Sminthidae family are called sminthids or birch mice. They are omnivores and primarily eat seeds, berries, and insects.[152] Sminthidae comprises sixteen extant species in a single genus.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Sicista
(birch mouse)

Brown birch mouse

Gray, 1827

Sixteen species
  • S. armenica (Armenian birch mouse)
  • S. betulina (Northern birch mouse, pictured)
  • S. caucasica (Caucasian birch mouse)
  • S. caudata (Long-tailed birch mouse)
  • S. cimlanica (Tsimlyansk birch mouse)
  • S. concolor (Chinese birch mouse)
  • S. kazbegica (Kazbeg birch mouse)
  • S. kluchorica (Kluchor birch mouse)
  • S. loriger (Nordmann's birch mouse)
  • S. napaea (Altai birch mouse)
  • S. pseudonapaea (Gray birch mouse)
  • S. severtzovi (Severtzov's birch mouse)
  • S. strandi (Strand's birch mouse)
  • S. subtilis (Southern birch mouse)
  • S. tianshanica (Tien Shan birch mouse)
  • S. trizona (Hungarian birch mouse)
Asia and Europe
Map of range
Size range: 4 cm (2 in) long, plus 8 cm (3 in) tail (Caucasian birch mouse) to 8 cm (3 in) long, plus 11 cm (4 in) tail (Altai birch mouse)[153]

Habitats: Inland wetlands, shrubland, forest, and grassland[154]

Diets: Seeds, berries, and insects[152]
Family Zapodidae

Members of the Zapodidae family are called zapodids or jumping mice. They are omnivores and eat a variety of vegetation and insects.[155] Zapodidae comprises five extant species in three genera.

Not assigned to a named subfamily – three genera
Name Authority and species Range Size and ecology
Eozapus Preble, 1899

One species
China
Map of range
Size: 7–10 cm (3–4 in) long, plus 11–15 cm (4–6 in) tail[156]

Habitats: Forest, shrubland, and grassland[157]

Diet: Vegetation and insects[155]
Napaeozapus

Brown mouse

Preble, 1899

One species
Eastern Canada and eastern United States
Map of range
Size: 8–10 cm (3–4 in) long, plus 11–16 cm (4–6 in) tail[156]

Habitats: Forest, shrubland, and inland wetlands[158]

Diet: Seeds, fungi, and insects, as well as fruit, nuts, and other vegetation[155]
Zapus
(meadow jumping mouse)

Brown mouse

Coues, 1875

Three species
Canada and United States
Map of range
Size range: 7 cm (3 in) long, plus 10 cm (4 in) tail (meadow jumping mouse) to 11 cm (4 in) long, plus 16 cm (6 in) tail (Pacific jumping mouse)[156]

Habitats: Forest, shrubland, grassland, and inland wetlands[159]

Diets: Seeds, as well as fungi, nuts, berries, fruit, and insects[155]

Superfamily Muroidea

Family Calomyscidae

Members of the Calomyscidae family are called calomyscids or brush-tailed mice. They are omnivores and primarily eat seeds, as well as flowers, leaves, and animal matter.[160] Calomyscidae comprises eight extant species in a single genus.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Calomyscus
(mouse-like hamster)

Brown hamster

Thomas, 1905

Eight species
  • C. bailwardi (Zagros Mountains mouse-like hamster)
  • C. baluchi (Baluchi mouse-like hamster)
  • C. elburzensis (Goodwin's brush-tailed mouse)
  • C. grandis (Noble mouse-like hamster)
  • C. hotsoni (Hotson's mouse-like hamster)
  • C. mystax (Great Balkhan mouse-like hamster)
  • C. tsolovi (Tsolov's mouse-like hamster)
  • C. urartensis (Urar mouse-like hamster, pictured)
Central Asia Size range: 6 cm (2 in) long, plus 5 cm (2 in) tail (Tsolov's mouse-like hamster) to 10 cm (4 in) long, plus 11 cm (4 in) tail (great Balkhan mouse-like hamster)[161]

Habitats: Shrubland, grassland and rocky areas[162]

Diets: Seeds, as well as flowers, leaves, and animal matter[160]
Family Cricetidae

Members of the Cricetidae family are called cricetids and include hamsters, voles, lemmings, muskrats, and New World rats and mice. They eat vegetation, seeds, and insects, and some also eat fungi, fish, and small animals.[163][164][165][166][167] Cricetidae comprises 694 extant species, the second-most of any mammal family, divided into 141 genera. These genera are divided into five subfamilies: Arvicolinae, containing 158 species of voles, lemmings, and muskrats; Cricetinae, containing 18 species of hamsters; Neotominae, containing 123 species of deer mice, packrats, and grasshopper mice; Sigmodontinae, containing 385 species of New World rats and mice; and Tylomyinae, containing 10 species of vesper and climbing rats. Additionally, one species in Neotominae and ten species Sigmodontinae in were driven extinct after 1500 due to the European colonization of the Americas and introduction of non-native rats, with some species surviving until the 1900s.[168][169]

Subfamily ArvicolinaeGray, 1821 – 30 genera
Name Authority and species Range Size and ecology
Alexandromys
(grass vole)

Gray vole

Ognev, 1914

Eleven species
  • A. evoronensis (Evorsk vole)
  • A. fortis (Reed vole, pictured)
  • A. kikuchii (Taiwan vole)
  • A. limnophilus (Lacustrine vole)
  • A. maximowiczii (Maximowicz's vole)
  • A. middendorffii (Middendorff's vole)
  • A. mongolicus (Mongolian vole)
  • A. montebelli (Japanese grass vole)
  • A. mujanensis (Muya Valley vole)
  • A. oeconomus (Tundra vole)
  • A. sachalinensis (Sakhalin vole)
Eastern and northern Asia, Europe, and northwestern North America Size range: 7 cm (3 in) long, plus 2 cm (0.8 in) tail (tundra vole) to 18 cm (7 in) long, plus 8 cm (3 in) tail (reed vole)[170]

Habitats: Forest, shrubland, inland wetlands, desert, rocky areas, and grassland[171]

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[163]
Alticola
(high mountain vole)

Gray and brown vole

Blanford, 1881

Twelve species
  • A. albicaudus (White-tailed mountain vole)
  • A. argentatus (Silver mountain vole, pictured)
  • A. barakshin (Gobi Altai mountain vole)
  • A. lemminus (Lemming vole)
  • A. macrotis (Large-eared vole)
  • A. montosa (Central Kashmir vole)
  • A. olchonensis (Lake Baikal mountain vole)
  • A. roylei (Royle's mountain vole)
  • A. semicanus (Mongolian silver vole)
  • A. stoliczkanus (Stolička's mountain vole)
  • A. strelzowi (Flat-headed vole)
  • A. tuvinicus (Tuva silver vole)
Asia Size range: 8 cm (3 in) long, plus 1 cm (0.4 in) tail (lemming vole) to 14 cm (6 in) long, plus 5 cm (2 in) tail (flat-headed vole)[170]

Habitats: Forest, rocky areas, shrubland, and grassland[172]

Diets: Stems and leaves[163]
Arborimus
(tree vole)

Brown vole

Taylor, 1915

Three species
  • A. albipes (White-footed vole)
  • A. longicaudus (Red tree vole, pictured)
  • A. pomo (Sonoma tree vole)
Western United States Size range: 9–11 cm (4 in) long, plus 6–9 cm (2–4 in) tail (red tree vole)[170]

Habitats: Forest and inland wetlands[173]

Diets: Conifer needles[163]
Arvicola
(water vole)

Brown vole

Lacépède, 1799

Three species
Europe and Asia Size range: 12–23 cm (5–9 in) long, plus 5–14 cm (2–6 in) tail (European water vole)[170]

Habitats: Forest, inland wetlands, and grassland[174]

Diets: Aquatic plants, herbs, grass, twigs, buds, roots, bulbs, and fruit[163]
Caryomys
(brownish vole)
Thomas, 1911

Two species
Central China Size range: 8–11 cm (3–4 in) long, plus 4–6 cm (2 in) tail (Ganzu vole)[170]

Habitats: Forest[175]

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[163]
Chionomys
(snow vole)

Brown vole

Miller, 1908

Three species
  • C. gud (Caucasian snow vole)
  • C. nivalis (European snow vole, pictured)
  • C. roberti (Robert's snow vole)
Europe and western Asia Size range: 10 cm (4 in) long, plus 4 cm (2 in) tail (European snow vole) to 16 cm (6 in) long, plus 11 cm (4 in) tail (Robert's snow vole)[170]

Habitats: Forest and rocky areas[176]

Diets: Grass, herbs, and seeds[163]
Clethrionomys
(bank vole)

Brown vole

Gistel, 1850

Five species
Europe, Asia, and North America Size range: 8–14 cm (3–6 in) long, plus 3–8 cm (1–3 in) tail (bank vole)[170]

Habitats: Forest, rocky areas, shrubland, and grassland[177]

Diets: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[163]
Craseomys
(grey-sided vole)

Brown vole

Miller, 1900

Six species
Northern Europe and northern and eastern Asia Size range: 7 cm (3 in) long, plus 4 cm (2 in) tail (Japanese red-backed vole) to 15 cm (6 in) long, plus 7 cm (3 in) tail (grey red-backed vole)[170]

Habitats: Forest, shrubland, inland wetlands, rocky areas, and grassland[178]

Diets: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[163]
Dicrostonyx
(collared lemming)

Gray and brown lemming

Gloger, 1841

Eight species
Russia and northern North America Size range: 8 cm (3 in) long, plus 1 cm (0.4 in) tail (Arctic lemming) to 16 cm (6 in) long, plus 3 cm (1 in) tail (northern collared lemming)[170]

Habitats: Rocky areas, shrubland, inland wetlands, and grassland[179]

Diets: Fruit, flowers, grass, sedges, buds, twigs, and bark[163]
Dinaromys Kretzoi, 1955

One species
Southern Europe
Map of range
Size: 13–16 cm (5–6 in) long, plus 8–11 cm (3–4 in) tail[170]

Habitats: Rocky areas[180]

Diet: Grass[163]
Ellobius
(mole vole)

Brown vole

Fischer von Waldheim, 1814

Five species
Eastern Europe and western and central Asia Size range: 8 cm (3 in) long, plus 0.5 cm (0 in) tail (northern mole vole) to 15 cm (6 in) long, plus 2 cm (0.8 in) tail (southern mole vole)[170]

Habitats: Desert, shrubland, and grassland[181]

Diets: Bulbs, tubers, and other underground plant parts[163]
Eolagurus
(steppe lemming)

Drawing of brown lemmings

Argiropulo, 1946

Two species
  • E. luteus (Yellow steppe lemming)
  • E. przewalskii (Przewalski's steppe lemming, pictured)
Central Asia Size range: 12 cm (5 in) long, plus 0.5 cm (0 in) tail (Przewalski's steppe lemming) to 16 cm (6 in) long, plus 3 cm (1 in) tail (yellow steppe lemming)[170]

Habitats: Shrubland and grassland[182]

Diets: Bulbs, seeds, and roots[163]
Eothenomys
(Père David's vole)

Drawing of brown vole

Miller, 1896

Eight species
  • E. cachinus (Kachin red-backed vole)
  • E. chinensis (Pratt's vole)
  • E. custos (Southwest China vole)
  • E. melanogaster (Père David's vole, pictured)
  • E. miletus (Yunnan red-backed vole)
  • E. olitor (Chaotung vole)
  • E. proditor (Yulungshan vole)
  • E. wardi (Ward's red-backed vole)
Eastern Asia Size range: 7 cm (3 in) long, plus 2 cm (0.8 in) tail (Père David's vole) to 14 cm (6 in) long, plus 7 cm (3 in) tail (Kachin red-backed vole)[170]

Habitats: Forest, rocky areas, shrubland, and grassland[183]

Diets: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[163]
Hyperacrius
(Kashmir vole)
Miller, 1896

Two species
  • H. fertilis (True's vole)
  • H. wynnei (Murree vole)
Northern Pakistan and western China Size range: 8 cm (3 in) long, plus 1 cm (0.4 in) tail (True's vole) to 14 cm (6 in) long, plus 5 cm (2 in) tail (Murree vole)[170]

Habitats: Forest, shrubland, and grassland[184]

Diets: Grass, stems, and roots[163]
Lagurus

Gray lemming

Gloger, 1841

One species
Eastern Europe and western, central, and eastern Asia Size: 7–13 cm (3–5 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[170]

Habitats: Shrubland and grassland[185]

Diet: Green vegetation, tubers, and bulbs[163]
Lasiopodomys
(narrow-headed steppe vole)

Brown vole

Lataste, 1887

Two species
  • L. brandtii (Brandt's vole)
  • L. mandarinus (Mandarin vole, pictured)
Eastern Asia Size range: 9 cm (4 in) long, plus 2 cm (0.8 in) tail (Mandarin vole) to 15 cm (6 in) long, plus 3 cm (1 in) tail (Brandt's vole)[170]

Habitats: Shrubland and grassland[186]

Diets: Grass, vegetation, and roots[163]
Lemmiscus

Gray vole

Thomas, 1912

One species
Western United States and western Canada
Map of range
Size: 9–12 cm (4–5 in) long, plus 1–3 cm (0–1 in) tail[170]

Habitats: Shrubland and grassland[187]

Diet: Green vegetation[163]
Lemmus
(lemming)

Brown and yellow lemming

Link, 1795

Five species
Northern Europe, northern Asia, and western North America Size range: 8 cm (3 in) long, plus 1 cm (0.4 in) tail (Amur lemming) to 16 cm (6 in) long, plus 2 cm (0.8 in) tail (West Siberian lemming)[170]

Habitats: Forest, shrubland, inland wetlands, and grassland[188]

Diets: Sedges, grass, bark, leaves, berries, lichens, and roots[163]
Microtus
(meadow vole)

Brown vole

Schrank, 1798

57 species
  • M. abbreviatus (Insular vole)
  • M. afghanus (Afghan vole)
  • M. agrestis (Short-tailed field vole)
  • M. anatolicus (Anatolian vole)
  • M. arvalis (Common vole)
  • M. brachycercus (Calabria pine vole)
  • M. breweri (Beach vole)
  • M. bucharensis (Bucharian vole)
  • M. cabrerae (Cabrera's vole)
  • M. californicus (California vole)
  • M. canicaudus (Gray-tailed vole)
  • M. chrotorrhinus (Rock vole)
  • M. daghestanicus (Daghestan pine vole)
  • M. dogramacii (Doğramaci's vole)
  • M. duodecimcostatus (Mediterranean pine vole)
  • M. felteni (Felten's vole)
  • M. gerbei (Gerbe's vole)
  • M. guatemalensis (Guatemalan vole)
  • M. guentheri (Günther's vole)
  • M. ilaeus (Tien Shan vole)
  • M. irani (Persian vole)
  • M. kermanensis (Kerman vole)
  • M. lavernedii (Mediterranean field vole)
  • M. liechtensteini (Liechtenstein's pine vole)
  • M. longicaudus (Long-tailed vole)
  • M. lusitanicus (Lusitanian pine vole)
  • M. majori (Major's pine vole)
  • M. mexicanus (Mexican vole)
  • M. miurus (Singing vole)
  • M. montanus (Montane vole)
  • M. multiplex (Alpine pine vole)
  • M. mystacinus (Caspian gray vole)
  • M. nebrodensis (Sicilian pine vole)
  • M. oaxacensis (Tarabundí vole)
  • M. obscurus (Altai vole)
  • M. ochrogaster (Prairie vole)
  • M. oregoni (Creeping vole)
  • M. paradoxus (Paradox vole)
  • M. pennsylvanicus (Eastern meadow vole)
  • M. pinetorum (Woodland vole, pictured)
  • M. qazvinensis (Qazvin vole)
  • M. quasiater (Jalapan pine vole)
  • M. richardsoni (North American water vole)
  • M. rossiaemeridionalis (East European grey vole)
  • M. rozianus (Portuguese field vole)
  • M. savii (Savi's pine vole)
  • M. schelkovnikovi (Schelkovnikov's pine vole)
  • M. schidlovskii (Schidlovsky's vole)
  • M. socialis (Social vole)
  • M. subterraneus (European pine vole)
  • M. tatricus (Tatra pine vole)
  • M. thomasi (Thomas's pine vole)
  • M. townsendii (Townsend's vole)
  • M. transcaspicus (Transcaspian vole)
  • M. umbrosus (Zempoaltépec vole)
  • M. xanthognathus (Taiga vole)
  • M. yuldaschi (Juniper vole)
North America, Central America, Europe, and Asia Size range: 6 cm (2 in) long, plus 1 cm (0.4 in) tail (Afghan vole) to 18 cm (7 in) long, plus 10 cm (4 in) tail (North American water vole)[170]

Habitats: Forest, intertidal marine, coastal marine, savanna, shrubland, desert, inland wetlands, rocky areas, and grassland[189]

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[163]
Myopus

Gray lemming

Miller, 1910

One species
Northern Europe and northern Asia Size: 9–12 cm (4–5 in) long, plus 1–2 cm (0.4–0.8 in) tail[170]

Habitats: Forest and inland wetlands[190]

Diet: Moss, stems, and bark[163]
Neodon
(mountain vole)

Brown vole

Horsfield, 1841

Six species
  • M. clarkei (Clarke's vole)
  • N. forresti (Forrest's mountain vole)
  • N. fuscus (Plateau vole, pictured)
  • N. irene (Chinese scrub vole)
  • N. linzhiensis (Linzhi mountain vole)
  • N. sikimensis (Sikkim mountain vole)
China and southeastern Asia Size range: 8 cm (3 in) long, plus 2 cm (0.8 in) tail (Chinese scrub vole) to 15 cm (6 in) long, plus 5 cm (2 in) tail (plateau vole)[170]

Habitats: Forest, shrubland, rocky areas, and grassland[191]

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[163]
Neofiber

Brown muskrat

True, 1884

One species
Southeastern United States
Map of range
Size: 18–21 cm (7–8 in) long, plus 9–17 cm (4–7 in) tail[170]

Habitats: Grassland and inland wetlands[192]

Diet: Maidencane, as well as rushes, sedges, sawgrass, and mangrove bark[163]
Ondatra

Brown muskrat

Link, 1795

One species
North America, Europe, and Asia
Map of range
Size: 27–30 cm (11–12 in) long, plus 20–25 cm (8–10 in) tail[170]

Habitats: Inland wetlands[193]

Diet: Cattails, bulrushes, grass, and other vegetation, as well as crabs, crayfish, mussels, and small fish[163]
Phaiomys

Drawing of brown vole

Blyth, 1863

One species
Western China Size: 9–13 cm (4–5 in) long, plus 2–4 cm (1–2 in) tail[170]

Habitats: Forest, grassland, rocky areas, and shrubland[194]

Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[163]
Phenacomys
(heather vole)

Drawing of brown voles

Merriam, 1889

Two species
Northwestern United States and Canada Size range: 8–12 cm (3–5 in) long, plus 2–4 cm (0.8–1.6 in) tail (eastern heather vole)[170]

Habitats: Forest, shrubland, and grassland[195]

Diets: Bark, buds, heaths, forbs, berries, and seeds[163]
Proedromys
(Duke of Bedford's vole)
Thomas, 1911

Two species
Central China Size range: 9 cm (4 in) long, plus 3 cm (1 in) tail (Duke of Bedford's vole) to 14 cm (6 in) long, plus 9 cm (4 in) tail (Liangshan vole)[170]

Habitats: Forest and grassland[196]

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[163]
Prometheomys Satunin, 1901

One species
Western Asia Size: 12–17 cm (5–7 in) long, plus 3–6 cm (1–2 in) tail[170]

Habitats: Grassland[197]

Diet: Green vegetation and roots[163]
Stenocranius
(narrow-headed vole)

Drawing of brown vole

Kaschtschenko, 1901

Two species
Northern Asia Size range: 8–15 cm (3–6 in) long, plus 1–5 cm (0.4–2.0 in) tail (narrow-headed vole)[170]

Habitats: Grassland[198]

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[163]
Synaptomys
(bog lemming)

Brown lemming

Baird, 1857

Two species
Canada and northern United States Size range: 9 cm (4 in) long, plus 1 cm (0.4 in) tail (northern bog lemming) to 14 cm (6 in) long, plus 3 cm (1 in) tail (southern bog lemming)[170]

Habitats: Forest, shrubland, inland wetlands, and grassland[199]

Diets: Green vegetation as well as slugs, snails, and other invertebrates[163]
Volemys
(Szechuan vole)
Zagorodnyuk, 1990

Two species
  • V. millicens (Szechuan vole)
  • V. musseri (Marie's vole)
Central China Size range: 8 cm (3 in) long, plus 4 cm (2 in) tail (Szechuan vole) to 13 cm (5 in) long, plus 8 cm (3 in) tail (Marie's vole)[170]

Habitats: Forest, rocky areas, and grassland[200]

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[163]
Subfamily CricetinaeWaldheim, 1817 – seven genera
Name Authority and species Range Size and ecology
Allocricetulus
(Mongolian hamster)
Argiropulo, 1932

Two species
  • A. curtatus (Mongolian hamster)
  • A. eversmanni (Eversmann's hamster)
East-central and central Asia Size range: 10 cm (4 in) long, plus 1 cm (0.4 in) tail (Mongolian hamster) to 14 cm (6 in) long, plus 4 cm (2 in) tail (Eversmann's hamster)[201]

Habitats: Grassland and shrubland[202]

Diets: Grains, beans, lentils, roots, and plant material, as well as insects and frogs[164]
Cansumys Allen, 1928

One species
Central China Size: 12–14 cm (5–6 in) long, plus 9–11 cm (4 in) tail[201]

Habitats: Forest[203]

Diet: Leaves and grasses[203]
Cricetulus
(ratlike hamster)

Gray hamster

A. Milne-Edwards, 1867

Six species
Asia and eastern Europe Size range: 7 cm (3 in) long, plus 1 cm (0.4 in) tail (Chinese striped hamster) to 15 cm (6 in) long, plus 4 cm (2 in) tail (grey dwarf hamster)[201]

Habitats: Grassland, forest, inland wetlands, desert, rocky areas, and shrubland[204]

Diets: Shoots, seeds, and insects[164]
Cricetus
(golden hamster)

Brown hamster

Leske, 1779

One species
Europe and western and central Asia
Map of range
Size: 16–32 cm (6–13 in) long, plus 2–7 cm (1–3 in) tail[201]

Habitats: Shrubland and grassland[205]

Diet: Grains, beans, lentils, roots, and plant material, as well as insects and frogs[164]
Mesocricetus

Brown hamster

Nehring, 1898

Four species
Western Asia and eastern Europe Size range: 12 cm (5 in) long, plus 1 cm (0.4 in) tail (golden hamster) to 22 cm (9 in) long, plus 2 cm (0.8 in) tail (Ciscaucasian hamster)[201]

Habitats: Grassland and shrubland[206]

Diets: Omnivorous, including a variety of vegetation, seeds, fruit, insects, and small vertebrates[164]
Phodopus
(dwarf hamster)

Brown hamster

Miller, 1910

Three species
East-central Asia and Kazakhstan and southern Russia Size range: 7 cm (3 in) long, plus 0.5 cm (0.2 in) tail (Roborovski dwarf hamster) to 11 cm (4 in) long, plus 2 cm (0.8 in) tail (Campbell's dwarf hamster)[201]

Habitats: Desert and shrubland[207]

Diets: Seeds and plant material[164]
Tscherskia

Brown hamster

Ogniov, 1914

One species
Eastern Asia Size: 14–22 cm (6–9 in) long, plus 6–11 cm (2–4 in) tail[201]

Habitats: Inland wetlands[208]

Diet: Seeds as well as shoots[164]
Subfamily NeotominaeMerriam, 1894 – sixteen genera
Name Authority and species Range Size and ecology
Baiomys
(New World pygmy mouse)

Gray mouse

True, 1894

Two species
  • B. musculus (Mexican pygmy mouse, pictured)
  • B. taylori (Northern pygmy mouse)
Southern United States, Mexico, and northern Central America Size range: 5 cm (2 in) long, plus 3 cm (1 in) tail (northern pygmy mouse) to 8 cm (3 in) long, plus 6 cm (2 in) tail (Mexican pygmy mouse)[168]

Habitats: Grassland, desert, forest, and shrubland[209]

Diets: Cactus stems and fruit, green vegetation, seeds, grass, and insects[165]
Habromys
(slender-tailed deer mouse)
Hooper & Musser, 1964

Six species
  • H. chinanteco (Chinanteco deer mouse)
  • H. delicatulus (Delicate deer mouse)
  • H. ixtlani (Ixtlán deer mouse)
  • H. lepturus (Zempoaltepec deer mouse)
  • H. lophurus (Crested-tailed deer mouse)
  • H. simulatus (Jico deer mouse)
Mexico and northern Central America Size range: 7 cm (3 in) long, plus 7 cm (3 in) tail (delicate deer mouse) to 14 cm (6 in) long, plus 15 cm (6 in) tail (Ixtlán deer mouse)[168]

Habitats: Forest[210]

Diets: Plant material and invertebrates[165]
Hodomys

Mouse jawbone

Merriam, 1894

One species
  • H. alleni (Allen's woodrat)
Southern Mexico Size: 16–22 cm (6–9 in) long, plus 14–22 cm (6–9 in) tail[168]

Habitats: Shrubland[211]

Diet: Plant material and invertebrates[165]
Isthmomys Hooper & Musser, 1964

Two species
  • I. flavidus (Yellow isthmus rat)
  • I. pirrensis (Mount Pirri isthmus rat)
Panama and northwestern Colombia Size range: 15–18 cm (6–7 in) long, plus 18–20 cm (7–8 in) tail (Mount Pirri isthmus rat)[168]

Habitats: Forest[212]

Diets: Plant material and invertebrates[165]
Megadontomys
(giant deer mouse)

Brown mouse

Merriam, 1898

Three species
  • M. cryophilus (Oaxaca giant deer mouse, pictured)
  • M. nelsoni (Nelson's giant deer mouse)
  • M. thomasi (Thomas's giant deer mouse)
Southern Mexico Size range: 13 cm (5 in) long, plus 17 cm (7 in) tail (Nelson's giant deer mouse) to 17 cm (7 in) long, plus 19 cm (7 in) tail (Thomas's giant deer mouse)[168]

Habitats: Forest[213]

Diets: Berries and seeds[213]
Nelsonia
(diminutive woodrat)
Merriam, 1897

Two species
  • N. goldmani (Goldman's diminutive woodrat)
  • N. neotomodon (Diminutive woodrat)
Central Mexico Size range: 11 cm (4 in) long, plus 10 cm (4 in) tail (diminutive woodrat) to 14 cm (6 in) long, plus 13 cm (5 in) tail (Goldman's diminutive woodrat)[168]

Habitats: Forest[214]

Diets: Conifer needles[165]
Neotoma
(pack rat)

Gray mouse

Say & Ord, 1825

20 species
North America and Central America Size range: 14 cm (6 in) long, plus 10 cm (4 in) tail (Arizona woodrat) to 25 cm (10 in) long, plus 21 cm (8 in) tail (big-eared woodrat)[168]

Habitats: Desert, shrubland, grassland, rocky areas, forest, caves, coastal marine, and inland wetlands[215]

Diets: Roots, stems, and leaves, as well as seeds and invertebrates[165]
Neotomodon

Brown mouse

Merriam, 1898

One species
Central Mexico Size: 9–13 cm (4–5 in) long, plus 7–11 cm (3–4 in) tail[168]

Habitats: Forest[216]

Diet: Plant material and invertebrates[165]
Ochrotomys

Drawing of brown mouse

Osgood, 1909

One species
Eastern United States Size: 5–12 cm (2–5 in) long, plus 5–10 cm (2–4 in) tail[168]

Habitats: Shrubland and forest[217]

Diet: Seeds[165]
Onychomys
(grasshopper mouse)

Brown and white mouse

Baird, 1857

Three species
United States and Mexico Size range: 8 cm (3 in) long, plus 4 cm (2 in) tail (Mearns's grasshopper mouse) to 13 cm (5 in) long, plus 7 cm (3 in) tail (northern grasshopper mouse)[168]

Habitats: Grassland, desert, and shrubland[218]

Diets: Insects, scorpions, and small vertebrates, as well as plant material[165]
Osgoodomys

Brown mouse

Hooper & Musser, 1964

One species
  • O. banderanus (Michoacan deer mouse)
Southwestern Mexico Size: 10–15 cm (4–6 in) long, plus 9–13 cm (4–5 in) tail[168]

Habitats: Forest[219]

Diet: Plant material and invertebrates[165]
Peromyscus
(deer mouse)

Brown mouse

Gloger, 1841

57 species
  • P. attwateri (Texas mouse)
  • P. aztecus (Aztec mouse)
  • P. beatae (Orizaba deer mouse)
  • P. boylii (Brush mouse)
  • P. bullatus (Perote mouse)
  • P. californicus (California deermouse)
  • P. caniceps (Burt's deer mouse)
  • P. crinitus (Canyon mouse)
  • P. dickeyi (Dickey's deer mouse)
  • P. difficilis (Zacatecan deer mouse)
  • P. eremicus (Cactus mouse)
  • P. eva (Eva's desert mouse)
  • P. fraterculus (Northern Baja deer mouse)
  • P. furvus (Blackish deer mouse)
  • P. gossypinus (Cotton mouse)
  • P. grandis (Big deer mouse)
  • P. gratus (Osgood's mouse)
  • P. guardia (Angel Island mouse)
  • P. guatemalensis (Guatemalan deer mouse)
  • P. gymnotis (Naked-eared deer mouse)
  • P. hooperi (Hooper's mouse)
  • P. hylocetes (Transvolcanic deer mouse)
  • P. interparietalis (San Lorenzo mouse)
  • P. keeni (Northwestern deer mouse)
  • P. leucopus (White-footed mouse)
  • P. levipes (Nimble-footed mouse)
  • P. madrensis (Tres Marías Island mouse)
  • P. maniculatus (Eastern deer mouse)
  • P. mayensis (Maya mouse)
  • P. megalops (Brown deer mouse)
  • P. mekisturus (Puebla deer mouse)
  • P. melanocarpus (Black-wristed deer mouse)
  • P. melanophrys (Plateau mouse)
  • P. melanotis (Black-eared mouse)
  • P. melanurus (Black-tailed mouse)
  • P. merriami (Mesquite mouse)
  • P. mexicanus (Mexican deer mouse)
  • P. nasutus (Northern rock mouse)
  • P. ochraventer (El Carrizo deer mouse)
  • P. pectoralis (White-ankled mouse)
  • P. pembertoni (Pemberton's deer mouse)
  • P. perfulvus (Tawny deer mouse)
  • P. polionotus (Oldfield mouse)
  • P. polius (Chihuahuan mouse)
  • P. pseudocrinitus (False canyon mouse)
  • P. sagax (La Palma deermouse)
  • P. schmidlyi (Schmidly's deermouse)
  • P. sejugis (Santa Cruz mouse)
  • P. simulus (Nayarit mouse)
  • P. slevini (Slevin's mouse, pictured)
  • P. spicilegus (Gleaning mouse)
  • P. stephani (San Esteban Island mouse)
  • P. stirtoni (Stirton's deer mouse)
  • P. truei (Pinyon mouse)
  • P. winkelmanni (Winkelmann's mouse)
  • P. yucatanicus (Yucatan deer mouse)
  • P. zarhynchus (Chiapan deer mouse)
North America and Central America Size range: 7 cm (3 in) long, plus 8 cm (3 in) tail (cactus mouse) to 15 cm (6 in) long, plus 16 cm (6 in) tail (brown deer mouse)[168]

Habitats: Desert, shrubland, grassland, rocky areas, forest, caves, intertidal marine, coastal marine, and inland wetlands[220]

Diets: A wide variety of plant material, fungi, insects, other invertebrates, and carrion, as well as worms, molluscs, and small vertebrates[165]
Podomys

Gray mouse

Osgood, 1909

One species
Southeastern United States Size: 9–12 cm (4–5 in) long, plus 8–11 cm (3–4 in) tail[168]

Habitats: Savanna, forest, grassland, and shrubland[221]

Diet: Seeds, nuts, fungi, vegetation, insects, and other invertebrates[165]
Reithrodontomys
(harvest mouse)

Brown mouse

Giglioli, 1873

21 species
  • R. brevirostris (Short-nosed harvest mouse)
  • R. burti (Sonoran harvest mouse)
  • R. chrysopsis (Volcano harvest mouse)
  • R. creper (Chiriqui harvest mouse)
  • R. darienensis (Darien harvest mouse)
  • R. fulvescens (Fulvous harvest mouse)
  • R. gracilis (Slender harvest mouse)
  • R. hirsutus (Hairy harvest mouse)
  • R. humulis (Eastern harvest mouse)
  • R. megalotis (Western harvest mouse, pictured)
  • R. mexicanus (Mexican harvest mouse)
  • R. microdon (Small-toothed harvest mouse)
  • R. montanus (Plains harvest mouse)
  • R. musseri (Small harvest mouse)
  • R. paradoxus (Nicaraguan harvest mouse)
  • R. raviventris (Salt marsh harvest mouse)
  • R. rodriguezi (Rodriguez's harvest mouse)
  • R. spectabilis (Cozumel harvest mouse)
  • R. sumichrasti (Sumichrast's harvest mouse)
  • R. tenuirostris (Narrow-nosed harvest mouse)
  • R. zacatecae (Zacatecas harvest mouse)
North America and Central America Size range: 5 cm (2 in) long, plus 5 cm (2 in) tail (plains harvest mouse) to 13 cm (5 in) long, plus 13 cm (5 in) tail (narrow-nosed harvest mouse)[168]

Habitats: Desert, shrubland, grassland, forest, intertidal marine, and inland wetlands[222]

Diets: Seeds and shoots, as well as insects[165]
Scotinomys
(singing mouse)

Drawing of brown mouse

Thomas, 1913

Two species
Southern Mexico and Central America Size range: 6 cm (2 in) long, plus 4 cm (2 in) tail (Alston's brown mouse) to 9 cm (4 in) long, plus 8 cm (3 in) tail (Chiriqui brown mouse)[168]

Habitats: Grassland and forest[223]

Diets: Insects[165]
Xenomys

Gray mouse

Merriam, 1892

One species
Southwestern Mexico Size: 15–17 cm (6–7 in) long, plus 14–17 cm (6–7 in) tail[168]

Habitats: Forest[224]

Diet: Plant material and invertebrates[165]
Subfamily SigmodontinaeWagner, 1843 – 87 genera
Name Authority and species Range Size and ecology
Abrawayaomys Cunha & Cruz, 1979

One species
Southeastern Brazil Size: 8–14 cm (3–6 in) long, plus 8–15 cm (3–6 in) tail[169]

Habitats: Forest[225]

Diet: Vegetation, seeds, and insects[166]
Abrothrix
(soft-haired mouse)

Brown and white mouse

Waterhouse, 1837

Eight species
  • A. andinus (Andean Altiplano mouse)
  • A. illuteus (Gray grass mouse)
  • A. jelskii (Jelski's Altiplano mouse, pictured)
  • A. lanosus (Woolly grass mouse)
  • A. longipilis (Long-haired grass mouse)
  • A. manni (Mann's grass mouse)
  • A. olivaceus (Olive grass mouse)
  • A. sanborni (Sanborn's grass mouse)
Western South America Size range: 6 cm (2 in) long, plus 4 cm (2 in) tail (Andean Altiplano mouse) to 14 cm (6 in) long, plus 10 cm (4 in) tail (long-haired grass mouse)[169]

Habitats: Rocky areas, inland wetlands, grassland, forest, and shrubland[226]

Diets: Insects, berries, seeds, and fungi[166]
Aegialomys
(Galápagos rice rat)

Gray rat

Weksler, Percequillo, & Voss, 2006

Two species
  • A. galapagoensis (Galápagos rice rat, pictured)
  • A. xanthaeolus (Yellowish rice rat)
Ecuador and western Peru Size range: 10–16 cm (4–6 in) long, plus 12–18 cm (5–7 in) tail (Galápagos rice rat)[169]

Habitats: Desert, forest, and shrubland[227]

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish[166]
Aepeomys
(montane mouse)
Thomas, 1898

Two species
  • A. lugens (Olive montane mouse)
  • A. reigi (Reig's montane mouse)
Western Venezuela Size range: 10–13 cm (4–5 in) long, plus 11–15 cm (4–6 in) tail (Reig's montane mouse)[169]

Habitats: Grassland, forest, and shrubland[228]

Diets: Vegetation, seeds, and insects[166]
Akodon
(grass mouse)

Brown mouse

Meyen, 1833

39 species
  • A. aerosus (Highland grass mouse)
  • A. affinis (Colombian grass mouse)
  • A. albiventer (White-bellied grass mouse)
  • A. azarae (Azara's grass mouse)
  • A. boliviensis (Bolivian grass mouse)
  • A. budini (Budin's grass mouse)
  • A. caenosus (Unicolored grass mouse)
  • A. cursor (Cursor grass mouse)
  • A. dayi (Day's grass mouse)
  • A. dolores (Dolorous grass mouse)
  • A. fumeus (Smoky grass mouse)
  • A. iniscatus (Intelligent grass mouse)
  • A. juninensis (Junín grass mouse)
  • A. kofordi (Koford's grass mouse)
  • A. lindberghi (Lindbergh's grass mouse)
  • A. lutescens (Altiplano grass mouse)
  • A. mimus (Thespian grass mouse)
  • A. molinae (Molina's grass mouse)
  • A. mollis (Soft grass mouse, pictured)
  • A. montensis (Montane grass mouse)
  • A. mystax (Caparaó grass mouse)
  • A. oenos (Monte grass mouse)
  • A. orophilus (El Dorado grass mouse)
  • A. paranaensis (Paraná grass mouse)
  • A. pervalens (Tarija akodont)
  • A. philipmyersi (Philip Myers's akodont)
  • A. polopi (Polop's grass mouse)
  • A. reigi (Reig's grass mouse)
  • A. sanctipaulensis (São Paulo grass mouse)
  • A. serrensis (Serra do Mar grass mouse)
  • A. siberiae (Cochabamba grass mouse)
  • A. simulator (White-throated grass mouse)
  • A. spegazzinii (Spegazzini's grass mouse)
  • A. subfuscus (Puno grass mouse)
  • A. surdus (Silent grass mouse)
  • A. sylvanus (Forest grass mouse)
  • A. toba (Chaco grass mouse)
  • A. torques (Cloud forest grass mouse)
  • A. varius (Variable grass mouse)
South America Size range: 6 cm (2 in) long, plus 4 cm (2 in) tail (Caparaó grass mouse) to 16 cm (6 in) long, plus 11 cm (4 in) tail (variable grass mouse)[169]

Habitats: Rocky areas, inland wetlands, savanna, grassland, forest, and shrubland[229]

Diets: Plants and invertebrates[166]
Amphinectomys Malygin, 1994

One species
  • A. savamis (Ucayali water rat)
Northern Peru Size: 18–19 cm (7 in) long, plus 17–21 cm (7–8 in) tail[169]

Habitats: Inland wetlands and forest[230]

Diet: Unknown[166]
Andalgalomys
(chaco mouse)
Williams & Mares, 1978

Two species
  • A. olrogi (Olrog's chaco mouse)
  • A. pearsoni (Pearson's chaco mouse)
Southern Bolivia, Paraguay, and Northern Argentina Size range: 7 cm (3 in) long, plus 8 cm (3 in) tail (Olrog's chaco mouse) to 13 cm (5 in) long, plus 14 cm (6 in) tail (Pearson's chaco mouse)[169]

Habitats: Grassland and shrubland[231]

Diets: Grass, grain, and mesquite fruit[166]
Andinomys Thomas, 1902

One species
  • A. edax (Andean mouse)
Western South America Size: 13–19 cm (5–7 in) long, plus 10–16 cm (4–6 in) tail[169]

Habitats: Grassland and shrubland[232]

Diet: Vegetation[166]
Anotomys , Genus Anotomys

One species
Ecuador and Colombia Size: 10–13 cm (4–5 in) long, plus 12–16 cm (5–6 in) tail[169]

Habitats: Inland wetlands, grassland, and forest[233]

Diet: Fish[166]
Auliscomys
(big-eared mouse)

Brown mouse

Thomas, 1906

Three species
  • A. boliviensis (Bolivian big-eared mouse, pictured)
  • A. pictus (Painted big-eared mouse)
  • A. sublimis (Andean big-eared mouse)
Western South America Size range: 9 cm (4 in) long, plus 4 cm (2 in) tail (Andean big-eared mouse) to 14 cm (6 in) long, plus 11 cm (4 in) tail (painted big-eared mouse)[169]

Habitats: Rocky areas, grassland, inland wetlands, and shrubland[234]

Diets: Seeds, vegetation, and lichen[166]
Bibimys
(crimson-nosed rat)

Brown mouse

Massoia, 1979

Three species
  • B. chacoensis (Chaco crimson-nosed rat)
  • B. labiosus (Large-lipped crimson-nosed rat, pictured)
  • B. torresi (Torres's crimson-nosed rat)
Paraguay, Argentina, and southern Brazil Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (large-lipped crimson-nosed rat) to 11 cm (4 in) long, plus 8 cm (3 in) tail (Torres's crimson-nosed rat)[169]

Habitats: Grassland, forest, inland wetlands, and shrubland[235]

Diets: Grass and seeds[166]
Blarinomys Thomas, 1896

One species
Northeastern Argentina and southern Brazil Size: 9–13 cm (4–5 in) long, plus 3–6 cm (1–2 in) tail[169]

Habitats: Forest[236]

Diet: Insects and worms[166]
Brucepattersonius
(brucie)
Hershkovitz, 1998

Seven species
  • B. griserufescens (Gray-bellied akodont)
  • B. guarani (Guaraní akodont)
  • B. igniventris (Red-bellied akodont)
  • B. iheringi (Ihering's akodont)
  • B. misionensis (Misiones akodont)
  • B. paradisus (Arroyo of Paradise brucie)
  • B. soricinus (Soricine brucie)
Northeastern Argentina and southern Brazil Size range: 8 cm (3 in) long, plus 8 cm (3 in) tail (Arroyo of Paradise brucie) to 13 cm (5 in) long, plus 10 cm (4 in) tail (red-bellied akodont)[169]

Habitats: Rocky areas and forest[237]

Diets: Vegetation, seeds, and insects[166]
Calomys
(vesper mouse)

Brown mouse

Waterhouse, 1837

Thirteen species
  • C. boliviae (Bolivian vesper mouse)
  • C. callidus (Crafty vesper mouse)
  • C. callosus (Large vesper mouse)
  • C. expulsus (Caatinga vesper mouse, pictured)
  • C. fecundus (Fecund vesper mouse)
  • C. hummelincki (Hummelinck's vesper mouse)
  • C. laucha (Small vesper mouse)
  • C. lepidus (Andean vesper mouse)
  • C. musculinus (Drylands vesper mouse)
  • C. sorellus (Peruvian vesper mouse)
  • C. tener (Delicate vesper mouse)
  • C. tocantinsi (Tocantins vesper mouse)
  • C. venustus (Córdoba vesper mouse)
South America Size range: 5 cm (2 in) long, plus 4 cm (2 in) tail (Hummelinck's vesper mouse) to 17 cm (7 in) long, plus 11 cm (4 in) tail (Córdoba vesper mouse)[169]

Habitats: Rocky areas, savanna, grassland, forest, and shrubland[238]

Diets: Vegetation and insects[166]
Casiomys
(Casio rice rat)

Brown rat

Voss, 2024

Six species
  • C. alfaroi (Alfaro's rice rat)
  • C. chapmani (Chapman's rice rat)
  • C. melanotis (Black-eared rice rat)
  • C. rhabdops (Striped rice rat)
  • C. rostratus (Long-nosed rice rat, pictured)
  • C. saturatior (Cloud Forest rice rat)
Mexico, Central America, and northeastern South America Size range: 8 cm (3 in) long, plus 9 cm (4 in) tail (black-eared rice rat) to 16 cm (6 in) long, plus 12 cm (5 in) tail (Chapman's rice rat)[169]

Habitats: Forest[239]

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish[166]
Cerradomys
(Cerrado rice rat)
Weksler, Percequillo, & Voss, 2006

Four species
  • C. maracajuensis (Maracaju rice rat)
  • C. marinhus (Marinho's rice rat)
  • C. scotti (Lindbergh's rice rat)
  • C. subflavus (Terraced rice rat)
South America Size range: 12–19 cm (5–7 in) long, plus 14–20 cm (6–8 in) tail (Lindbergh's rice rat)[169]

Habitats: Inland wetlands, savanna, grassland, forest, and shrubland[240]

Diets: Vegetation, seeds, and insects[166]
Chelemys
(long-clawed mouse)

Drawing of brown and white mice

Thomas, 1903

Two species
  • C. macronyx (Andean long-clawed mouse)
  • C. megalonyx (Large long-clawed mouse, pictured)
Chile and southwestern Argentina Size range: 10–15 cm (4–6 in) long, plus 4–7 cm (2–3 in) tail (Andean long-clawed mouse)[169]

Habitats: Grassland, forest, and shrubland[241]

Diets: Arthropods, vegetation, and fungi[166]
Chibchanomys
(water mouse)
Voss, 1988

Two species
Northwestern South America Size range: 10 cm (4 in) long, plus 10 cm (4 in) tail (Las Cajas water mouse) to 13 cm (5 in) long, plus 14 cm (6 in) tail (Chibchan water mouse)[169]

Habitats: Grassland, forest, and inland wetlands[242]

Diets: Aquatic invertebrates and small animals[166]
Chilomys Thomas, 1897

One species
Northwestern South America Size: 8–10 cm (3–4 in) long, plus 10–13 cm (4–5 in) tail[169]

Habitats: Forest[243]

Diet: Omnivorous[243]
Chinchillula Thomas, 1898

One species
  • C. sahamae (Altiplano chinchilla mouse)
Western South America
Map of range
Size: 15–17 cm (6–7 in) long, plus 9–11 cm (4 in) tail[169]

Habitats: Shrubland and rocky areas[244]

Diet: Vegetation[166]
Delomys
(Atlantic Forest rat)

Brown rat

Thomas, 1917

Three species
  • D. collinus (Montane Atlantic Forest rat)
  • D. dorsalis (Striped Atlantic Forest rat, pictured)
  • D. sublineatus (Pallid Atlantic Forest rat)
Southeastern South America Size range: 10 cm (4 in) long, plus 10 cm (4 in) tail (montane Atlantic Forest rat) to 15 cm (6 in) long, plus 13 cm (5 in) tail (pallid Atlantic Forest rat)[169]

Habitats: Grassland and forest[245]

Diets: Vegetation, seeds, and insects[166]
Deltamys

Brown mouse

Thomas, 1917

One species
Southeastern South America Size: About 10 cm (4 in)[169]

Habitats: Inland wetlands and grassland[246]

Diet: Plants and invertebrates[166]
Drymoreomys

Brown rat

Percequillo, Weksler, & Costa, 2011

One species
Southeastern Brazil
Map of range
Size: 11–15 cm (4–6 in) long, plus 14–18 cm (6–7 in) tail[169]

Habitats: Forest[247]

Diet: Vegetation, seeds, and insects[166]
Eligmodontia
(gerbil mouse)
F. Cuvier, 1837

Four species
  • E. moreni (Monte gerbil mouse)
  • E. morgani (Morgan's gerbil mouse)
  • E. puerulus (Andean gerbil mouse)
  • E. typus (Lowland gerbil mouse)
Western South America Size range: 6–10 cm (2–4 in) long, plus 5–10 cm (2–4 in) tail (Andean gerbil mouse)[169]

Habitats: Desert, grassland, and shrubland[248]

Diets: Grain, vegetation, and insects[166]
Eremoryzomys Weksler, Percequillo, & Voss, 2006

One species
Northern Peru and southern Ecuador
Map of range
Size: 14–18 cm (6–7 in) long, plus 16–21 cm (6–8 in) tail[169]

Habitats: Forest[249]

Diet: Vegetation, seeds, and insects[166]
Euneomys
(chinchilla mouse)

Brown mouse

Coues, 1874

Four species
  • E. chinchilloides (Patagonian chinchilla mouse, pictured)
  • E. fossor (Burrowing chinchilla mouse)
  • E. mordax (Biting chinchilla mouse)
  • E. petersoni (Peterson's chinchilla mouse)
Chile and Argentina Size range: 7–16 cm (3–6 in) long, plus 4–9 cm (2–4 in) tail (Patagonian chinchilla mouse)[169]

Habitats: Rocky areas, grassland, forest, and shrubland[250]

Diets: Vegetation[166]
Euryoryzomys
(broad rice rat)

Brown rat

Weksler, Percequillo, & Voss, 2006

Six species
South America Size range: 9–18 cm (4–7 in) long, plus 9–19 cm (4–7 in) tail (Russet rice rat)[169]

Habitats: Savanna and forest[251]

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish[166]
Galenomys Thomas, 1916

One species
  • G. garleppi (Garlepp's mouse)
Western South America Size: 10–14 cm (4–6 in) long, plus 3–5 cm (1–2 in) tail[169]

Habitats: Grassland[252]

Diet: Seeds, vegetation, and lichen[166]
Geoxus
(long-clawed mole mouse)

Brown and white mouse

Thomas, 1919

Two species
  • G. annectens (Pearson's long-clawed akodont)
  • G. valdivianus (Long-clawed mole mouse, pictured)
Chile and southern Argentina Size range: 9 cm (4 in) long, plus 3 cm (1 in) tail (long-clawed mole mouse) to 13 cm (5 in) long, plus 9 cm (4 in) tail (Pearson's long-clawed akodont)[169]

Habitats: Forest and shrubland[253]

Diets: Worms, insects, and other arthropods[166]
Graomys
(gray leaf-eared mouse)

Gray mouse

Thomas, 1916

Four species
  • G. chacoensis (Central leaf-eared mouse)
  • G. domorum (Pale leaf-eared mouse)
  • G. edithae (Edith's leaf-eared mouse)
  • G. griseoflavus (Gray leaf-eared mouse, pictured)
Central and southern South America Size range: 10 cm (4 in) long, plus 14 cm (6 in) tail (pale leaf-eared mouse) to 17 cm (7 in) long, plus 18 cm (7 in) tail (gray leaf-eared mouse)[169]

Habitats: Grassland, forest, and shrubland[254]

Diets: Grass, grain, and mesquite fruit[166]
Gyldenstolpia

Drawing of gray rat

Pardiñas, D'Elía, & Teta, 2009

One species
  • G. fronto (Fossorial giant rat)
Southern Paraguay and northern Argentina Size: About 22 cm (9 in)[169]

Habitats: Inland wetlands and savanna[255]

Diet: Vegetation, seeds, and insects[166]
Handleyomys
(Handley's rice rat)
Voss, Gómez-Laverde, & Pacheco, 2002

Two species
  • H. fuscatus (Dusky-footed Handley's mouse)
  • H. intectus (Colombian rice rat)
Northwestern Colombia Size range: 8 cm (3 in) long, plus 7 cm (3 in) tail (Colombian rice rat) to 13 cm (5 in) long, plus 11 cm (4 in) tail (dusky-footed Handley's mouse)[169]

Habitats: Forest[256]

Diets: Vegetation, seeds, and insects[166]
Holochilus
(marsh rat)

Brown rat

Brandt, 1835

Three species
South America Size range: 13 cm (5 in) long, plus 12 cm (5 in) tail (Amazonian marsh rat) to 24 cm (9 in) long, plus 24 cm (9 in) tail (Brazilian marsh rat)[169]

Habitats: Grassland, forest, inland wetlands, and shrubland[257]

Diets: Marsh plants and molluscs[166]
Hylaeamys
(forest rice rat)

Brown rat

Weksler, Percequillo, & Voss, 2006

Eight species
  • H. acritus (Bolivian rice rat)
  • H. laticeps (Atlantic forest oryzomys)
  • H. megacephalus (Azara's broad-headed oryzomys, pictured)
  • H. oniscus (Sowbug rice rat)
  • H. perenensis (Western Amazonian rice rat)
  • H. seuanezi (Atlantic Forest rice rat)
  • H. tatei (Tate's rice rat)
  • H. yunganus (Yungas rice rat)
South America Size range: 8 cm (3 in) long, plus 9 cm (4 in) tail (Azara's broad-headed oryzomys) to 18 cm (7 in) long, plus 16 cm (6 in) tail (Atlantic Forest rice rat)[169]

Habitats: Inland wetlands, savanna, grassland, and forest[258]

Diets: Vegetation, seeds, and insects[166]
Ichthyomys
(crab-eating rat)

Drawing of brown and white rat

Thomas, 1893

Four species
  • I. hydrobates (Crab-eating rat, pictured)
  • I. pittieri (Pittier's crab-eating rat)
  • I. stolzmanni (Stolzmann's crab-eating rat)
  • I. tweedii (Tweedy's crab-eating rat)
Northwestern South America Size range: 9 cm (4 in) long, plus 8 cm (3 in) tail (Pittier's crab-eating rat) to 20 cm (8 in) long, plus 16 cm (6 in) tail (Tweedy's crab-eating rat)[169]

Habitats: Forest and inland wetlands[259]

Diets: Fish, crabs, and aquatic insects[166]
Irenomys Thomas, 1919

One species
Southern Chile and southwestern Argentina Size: 10–13 cm (4–5 in) long, plus 16–20 cm (6–8 in) tail[169]

Habitats: Shrubland and forest[260]

Diet: Fruit, seeds, and vegetation[166]
Juliomys
(Atlantic Forest tree mouse)

Brown mouse

González, 2000

Two species
  • J. pictipes (Lesser Wilfred's mouse)
  • J. rimofrons (Montane red-rumped tree mouse, pictured)
Southern Brazil and northeastern Argentina Size range: 8 cm (3 in) long, plus 9 cm (4 in) tail (montane red-rumped tree mouse) to 11 cm (4 in) long, plus 12 cm (5 in) tail (lesser Wilfred's mouse)[169]

Habitats: Forest[261]

Diets: Vegetation, seeds, and insects[166]
Juscelinomys
(Brazilian burrowing mouse)

Brown mouse

Oliveira, 1965

Two species
  • J. candango (Candango mouse, pictured)
  • J. huanchacae (Huanchaca mouse)
Northeastern Bolivia and Brazil Size range: 13–20 cm (5–8 in) long, plus 8–10 cm (3–4 in) tail (Huanchaca mouse)[169]

Habitats: Savanna, and grassland[262]

Diets: Vegetation and insects[166]
Kunsia

Gray rat

Hershkovitz, 1966

One species
Northern Bolivia and western Brazil Size: 18–29 cm (7–11 in) long, plus 14–20 cm (6–8 in) tail[169]

Habitats: Grassland[263]

Diet: Roots and grass[263]
Lenoxus Thomas, 1909

One species
Southern Peru and western Bolivia Size: 11–17 cm (4–7 in) long, plus 13–19 cm (5–7 in) tail[169]

Habitats: Forest[264]

Diet: Vegetation, seeds, and insects[166]
Loxodontomys
(big-eared mouse)

Drawing of brown mouse

Osgood, 1947

Two species
  • L. micropus (Southern big-eared mouse, pictured)
  • L. pikumche (Pikumche pericote)
Chile and southwestern Argentina Size range: 9–15 cm (4–6 in) long, plus 8–12 cm (3–5 in) tail (Pikumche pericote)[169]

Habitats: Shrubland[265]

Diets: Vegetation, fungi, and flowers[166]
Lundomys Voss & Carleton, 1993

One species
Uruguay and southern Brazil Size: 17–24 cm (7–9 in) long, plus 19–29 cm (7–11 in) tail[169]

Habitats: Inland wetlands, forest, and grassland[266]

Diet: Vegetation[166]
Megalomys 
(pilorie)

Brown mouse

Trouessart, 1881

Two species
Martinique and St. Lucia Size range: Unknown[169]

Habitats: Unknown[267]

Diets: Unknown[166]
Megaoryzomys 

Rat skull pieces

Lenglet & Coppois, 1979

One species
  • M. curioi (Galápagos giant rat)
Santa Cruz island of the Galápagos Islands Size: Unknown[169]

Habitats: Shrubland[268]

Diet: Unknown[166]
Melanomys
(dark rice rat)

Brown rat

Thomas, 1902

Three species
  • M. caliginosus (Dusky rice rat, pictured)
  • M. robustulus (Robust dark rice rat)
  • M. zunigae (Zuniga's dark rice rat)
Central America and northwestern South America Size range: 10–14 cm (4–6 in) long, plus 7–12 cm (3–5 in) tail (dusky rice rat)[169]

Habitats: Desert, forest, and shrubland[269]

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish[166]
Microakodontomys

Brown rats

Hershkovitz, 1993

One species
  • M. transitorius (Transitional colilargo)
Central Brazil Size: About 7 cm (3 in)[169]

Habitats: Shrubland and grassland[270]

Diet: Vegetation, seeds, and insects[166]
Microryzomys
(colilargo)
Thomas, 1917

Two species
Northwestern South America Size range: 6–10 cm (2–4 in) long, plus 11–13 cm (4–5 in) tail (montane colilargo)[169]

Habitats: Rocky areas, grassland, and forest[271]

Diets: Seeds and vegetation[271]
Mindomys Weksler, Percequillo, & Voss, 2006

One species
Ecuador
Map of range
Size: 17–29 cm (7–11 in) long, plus 22–25 cm (9–10 in) tail[169]

Habitats: Forest[272]

Diet: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish[166]
Neacomys
(bristly mouse)

Brown mouse

Thomas, 1900

Eight species
  • N. dubosti (Dubost's bristly mouse)
  • N. guianae (Guiana bristly mouse)
  • N. minutus (Small bristly mouse)
  • N. musseri (Musser's bristly mouse)
  • N. paracou (Paracou bristly mouse)
  • N. pictus (Painted bristly mouse)
  • N. spinosus (Common bristly mouse, pictured)
  • N. tenuipes (Narrow-footed bristly mouse)
Eastern Panama and South America Size range: 6 cm (2 in) long, plus 7 cm (3 in) tail (Dubost's bristly mouse) to 11 cm (4 in) long, plus 13 cm (5 in) tail (common bristly mouse)[169]

Habitats: Forest[273]

Diets: Seeds, insects, and fruit[273]
Necromys
(bolo mouse)

Brown mouse

Ameghino, 1889

Nine species
  • N. amoenus (Pleasant bolo mouse)
  • N. benefactus (Argentine bolo mouse)
  • N. lactens (Rufous-bellied bolo mouse)
  • N. lasiurus (Hairy-tailed bolo mouse, pictured)
  • N. lenguarum (Paraguayan bolo mouse)
  • N. obscurus (Dark bolo mouse)
  • N. punctulatus (Spotted bolo mouse)
  • N. temchuki (Temchuk's bolo mouse)
  • N. urichi (Northern grass mouse)
South America Size range: 9 cm (4 in) long, plus 6 cm (2 in) tail (pleasant bolo mouse) to 15 cm (6 in) long, plus 11 cm (4 in) tail (northern grass mouse)[169]

Habitats: Inland wetlands, savanna, grassland, forest, and shrubland[274]

Diets: Arthropods[166]
Nectomys
(water rat)

Brown rat

Peters, 1861

Five species
  • N. apicalis (Western Amazonian water rat)
  • N. magdalenae (Magdalena water rat)
  • N. palmipes (Trinidad water rat)
  • N. rattus (Common water rat)
  • N. squamipes (South American water rat, pictured)
South America and Trinidad and Tobago Size range: 11 cm (4 in) long, plus 12 cm (5 in) tail (South American water rat) to 29 cm (11 in) long, plus 25 cm (10 in) tail (common water rat)[169]

Habitats: Inland wetlands, savanna, grassland, forest, and shrubland[275]

Diets: Vegetation, insects, tadpoles, and small fish[166]
Neomicroxus
(small grass mouse)
Alvarado-Serrano & D'Elía, 2013

Two species
  • N. bogotensis (Bogotá grass mouse)
  • N. latebricola (Ecuadorian grass mouse)
Northwestern South America Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (Bogotá grass mouse) to 11 cm (4 in) long, plus 10 cm (4 in) tail (Ecuadorian grass mouse)[169]

Habitats: Grassland, forest, and shrubland[276]

Diets: Vegetation, seeds, and insects[166]
Neotomys

Brown rat

Thomas, 1894

One species
  • N. ebriosus (Andean swamp rat)
Western South America Size: 9–15 cm (4–6 in) long, plus 6–9 cm (2–4 in) tail[169]

Habitats: Inland wetlands, shrubland, and grassland[277]

Diet: Vegetation, seeds, and insects[166]
Nephelomys
(misty rice rat)

Brown rat

Weksler, Percequillo, & Voss, 2006

Seven species
Central America and Northwestern South America Size range: 10 cm (4 in) long, plus 13 cm (5 in) tail (Tomes's rice rat) to 18 cm (7 in) long, plus 20 cm (8 in) tail (Ecuadorian rice rat)[169]

Habitats: Grassland and forest[278]

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish[166]
Nesoryzomys
(Galápagos mouse)
Heller, 1904

Five species
  • N. darwini (Darwin's Galápagos mouse)
  • N. fernandinae (Fernandina Galápagos mouse)
  • N. indefessus (Indefatigable Galápagos mouse)
  • N. narboroughi (Large Fernandina Galápagos mouse)
  • N. swarthi (Santiago Galápagos mouse)
Galápagos Islands Size range: 10 cm (4 in) long, plus 7 cm (3 in) tail (Fernandina Galápagos mouse) to 19 cm (7 in) long, plus 17 cm (7 in) tail (Santiago Galápagos mouse)[169]

Habitats: Grassland, forest, and shrubland[279]

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish[166]
Neusticomys
(fish-eating rat)

Gray rat

Anthony, 1921

Six species
  • N. ferreirai (Ferreira's fish-eating rat)
  • N. monticolus (Montane fish-eating rat, pictured)
  • N. mussoi (Peruvian fish-eating rat)
  • N. oyapocki (Oyapock's fish-eating rat)
  • N. peruviensis (Peruvian fish-eating rat)
  • N. venezuelae (Venezuelan fish-eating rat)
Northern South America Size range: 9–21 cm (4–8 in) long, plus 8–12 cm (3–5 in) tail (montane fish-eating rat)[169]

Habitats: Savanna, forest, and inland wetlands[280]

Diets: Aquatic invertebrates[166]
Noronhomys Olson & Carleton, 1999

One species
Fernando de Noronha islands northeast of Brazil
Map of range
Size: Unknown[169]

Habitats: Shrubland[281]

Diet: Vegetation, seeds, and insects[166]
Notiomys Thomas, 1890

One species
Southern Argentina Size: 8–12 cm (3–5 in) long, plus 3–5 cm (1–2 in) tail[169]

Habitats: Shrubland, grassland, and rocky areas[282]

Diet: Insects and seeds[166]
Oecomys
(arboreal rice rat)
Thomas, 1906

Fifteen species
  • O. auyantepui (North Amazonian arboreal rice rat)
  • O. bicolor (Bicolored arboreal rice rat)
  • O. catherinae (Atlantic Forest arboreal rice rat)
  • O. cleberi (Cleber's arboreal rice rat)
  • O. concolor (Unicolored arboreal rice rat)
  • O. flavicans (Yellow arboreal rice rat)
  • O. mamorae (Mamore arboreal rice rat)
  • O. paricola (Brazilian arboreal rice rat)
  • O. phaeotis (Dusky arboreal rice rat)
  • O. rex (King arboreal rice rat)
  • O. roberti (Robert's arboreal rice rat)
  • O. rutilus (Red arboreal rice rat)
  • O. speciosus (Venezuelan arboreal rice rat)
  • O. superans (Foothill arboreal rice rat)
  • O. trinitatis (Trinidad arboreal rice rat)
South America Size range: 7 cm (3 in) long, plus 8 cm (3 in) tail (red arboreal rice rat) to 17 cm (7 in) long, plus 19 cm (7 in) tail (foothill arboreal rice rat)[169]

Habitats: Inland wetlands, savanna, grassland, forest, and shrubland[283]

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish[166]
Oligoryzomys
(pygmy rice rat)

Brown rat

Bangs, 1900

20 species
Central America and South America Size range: 7–12 cm (3–5 in) long, plus 10–15 cm (4–6 in) tail (black-footed pygmy rice rat)[169]

Habitats: Inland wetlands, savanna, grassland, forest, desert, and shrubland[284]

Diets: Seeds, fruit, and insects[166]
Oreoryzomys

Brown and white rat

Weksler, Percequillo, & Voss, 2006

One species
  • O. balneator (Peruvian rice rat)
Ecuador and northwestern Peru Size: 7–10 cm (3–4 in) long, plus 9–12 cm (4–5 in) tail[169]

Habitats: Forest[285]

Diet: Vegetation, seeds, and insects[166]
Oryzomys
(rice rat)

Gray rat

Baird, 1857

Six species
North America, Central America, South America, and Jamaica
Map of range
Size range: 9 cm (4 in) long, plus 10 cm (4 in) tail (Coues's rice rat) to 16 cm (6 in) long, plus 14 cm (6 in) tail (Gorgas's rice rat)[169]

Habitats: Inland wetlands, grassland, forest, intertidal marine, and shrubland[286]

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish[166]
Oxymycterus
(hocicudo)

Brown rat

Waterhouse, 1837

Seventeen species
  • O. akodontius (Argentine hocicudo)
  • O. amazonicus (Amazonian hocicudo)
  • O. angularis (Angular hocicudo)
  • O. caparoae (Caparaó hocicudo)
  • O. dasytrichus (Atlantic Forest hocicudo, pictured)
  • O. delator (Spy hocicudo)
  • O. hiska (Small hocicudo)
  • O. hispidus (Hispid hocicudo)
  • O. hucucha (Quechuan hocicudo)
  • O. inca (Incan hocicudo)
  • O. josei (Cook's hocicudo)
  • O. nasutus (Long-nosed hocicudo)
  • O. paramensis (Paramo hocicudo)
  • O. quaestor (Quaestor hocicudo)
  • O. roberti (Robert's hocicudo)
  • O. rufus (Red hocicudo)
  • O. wayku (Ravine hocicudo)
South America Size range: 8 cm (3 in) long, plus 10 cm (4 in) tail (Robert's hocicudo) to 20 cm (8 in) long, plus 16 cm (6 in) tail (angular hocicudo)[169]

Habitats: Inland wetlands, savanna, grassland, forest, and shrubland[287]

Diets: Insects, as well as other invertebrates and vegetation[166]
Phaenomys

Brown rat

Thomas, 1917

One species
  • P. ferrugineus (Rio de Janeiro arboreal rat)
Southeastern Brazil Size: 14–17 cm (6–7 in) long, plus 18–20 cm (7–8 in) tail[169]

Habitats: Forest[288]

Diet: Vegetation, seeds, and insects[166]
Phyllotis
(leaf-eared mouse)

Brown and white mouse

Waterhouse, 1837

Sixteen species
  • P. alisosiensis (Los Alisos leaf-eared mouse)
  • P. amicus (Friendly leaf-eared mouse)
  • P. andium (Andean leaf-eared mouse)
  • P. anitae (Anita's leaf-eared mouse)
  • P. bonariensis (Buenos Aires leaf-eared mouse)
  • P. caprinus (Capricorn leaf-eared mouse)
  • P. darwini (Darwin's leaf-eared mouse, pictured)
  • P. definitus (Definitive leaf-eared mouse)
  • P. gerbillus (Gerbil leaf-eared mouse)
  • P. haggardi (Haggard's leaf-eared mouse)
  • P. limatus (Lima leaf-eared mouse)
  • P. magister (Master leaf-eared mouse)
  • P. osgoodi (Osgood's leaf-eared mouse)
  • P. osilae (Bunchgrass leaf-eared mouse)
  • P. wolffsohni (Wolffsohn's leaf-eared mouse)
  • P. xanthopygus (Yellow-rumped leaf-eared mouse)
Western and southern South America Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (gerbil leaf-eared mouse) to 16 cm (6 in) long, plus 15 cm (6 in) tail (Buenos Aires leaf-eared mouse)[169]

Habitats: Rocky areas, inland wetlands, grassland, forest, desert, and shrubland[289]

Diets: Seeds, vegetation, and lichen[166]
Podoxymys Anthony, 1929

One species
  • P. roraimae (Roraima mouse)
Northern South America Size: 7–11 cm (3–4 in) long, plus 7–11 cm (3–4 in) tail[169]

Habitats: Forest[290]

Diet: Vegetation, seeds, and insects[166]
Pseudoryzomys Hershkovitz, 1962

One species
Central and eastern South America
Map of range
Size: 9–13 cm (4–5 in) long, plus 10–14 cm (4–6 in) tail[169]

Habitats: Grassland, savanna, and shrubland[291]

Diet: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish[166]
Punomys
(puna mouse)
Osgood, 1943

Two species
  • P. kofordi (Eastern puna mouse)
  • P. lemminus (Puna mouse)
Western South America Size range: 12–17 cm (5–7 in) long, plus 6–9 cm (2–4 in) tail (eastern puna mouse)[169]

Habitats: Rocky areas, grassland, and shrubland[292]

Diets: Twigs from herbs[166]
Reithrodon
(bunny rat)

Brown rat

Waterhouse, 1837

Two species
  • R. auritus (Bunny rat, pictured)
  • R. typicus (Naked-soled conyrat)
Southeastern South America Size range: 12–15 cm (5–6 in) long, plus 7–10 cm (3–4 in) tail (bunny rat)[169]

Habitats: Grassland and shrubland[293]

Diets: Grass, rhizomes, and roots[166]
Rhagomys
(arboreal mouse)
Thomas, 1917

Two species
  • R. longilingua (Long-tongued arboreal mouse)
  • R. rufescens (Brazilian arboreal mouse)
Southern South America Size range: 8 cm (3 in) long, plus 8 cm (3 in) tail (Brazilian arboreal mouse) to 11 cm (4 in) long, plus 11 cm (4 in) tail (long-tongued arboreal mouse)[169]

Habitats: Forest[294]

Diets: Vegetation, seeds, and insects[166]
Rheomys
(water mouse)

Brown and white rat head

Thomas, 1906

Four species
  • R. mexicanus (Mexican water mouse)
  • R. raptor (Goldman's water mouse)
  • R. thomasi (Thomas's water mouse)
  • R. underwoodi (Underwood's water mouse, pictured)
Southern Mexico and Central America Size range: 9 cm (4 in) long, plus 8 cm (3 in) tail (Goldman's water mouse) to 15 cm (6 in) long, plus 18 cm (7 in) tail (Mexican water mouse)[169]

Habitats: Forest and inland wetlands[295]

Diets: Fish, snails, and aquatic insects[166]
Rhipidomys
(climbing mouse)

Brown and white rat

Tschudi, 1845

22 species
  • R. albujai (Albuja's climbing rat)
  • R. austrinus (Southern climbing mouse)
  • R. cariri (Cariri climbing mouse)
  • R. caucensis (Cauca climbing mouse, pictured)
  • R. couesi (Coues's climbing mouse)
  • R. emiliae (Eastern Amazon climbing mouse)
  • R. fulviventer (Buff-bellied climbing mouse)
  • R. gardneri (Gardner's climbing mouse)
  • R. ipukensis (Ipuca climbing rat)
  • R. itoan (Sky climbing rat)
  • R. latimanus (Broad-footed climbing mouse)
  • R. leucodactylus (White-footed climbing mouse)
  • R. macconnelli (MacConnell's climbing mouse)
  • R. macrurus (Cerrado climbing mouse)
  • R. mastacalis (Atlantic Forest climbing mouse)
  • R. modicus (Peruvian climbing mouse)
  • R. nitela (Splendid climbing mouse)
  • R. ochrogaster (Yellow-bellied climbing mouse)
  • R. tribei (Tribe's climbing rat)
  • R. venezuelae (Venezuelan climbing mouse)
  • R. venustus (Charming climbing mouse)
  • R. wetzeli (Wetzel's climbing mouse)
Panama and South America Size range: 6 cm (2 in) long, plus 8 cm (3 in) tail (Wetzel's climbing mouse) to 21 cm (8 in) long, plus 20 cm (8 in) tail (Coues's climbing mouse)[169]

Habitats: Rocky areas, savanna, grassland, and forest[296]

Diets: Vegetation, seeds, and insects[166]
Salinomys Braun & Mares, 1995

One species
  • S. delicatus (Delicate salt flat mouse)
Northern Argentina Size: 6–9 cm (2–4 in) long, plus 9–13 cm (4–5 in) tail[169]

Habitats: Shrubland and inland wetlands[297]

Diet: Seeds, arthropods, and vegetation[297]
Scapteromys
(swamp rat)

Gray rat

Waterhouse, 1837

Two species
  • S. aquaticus (Argentine swamp rat, pictured)
  • S. tumidus (Waterhouse's swamp rat)
Southern South America Size range: 12–25 cm (5–10 in) long, plus 10–15 cm (4–6 in) tail (Argentine swamp rat)[169]

Habitats: Grassland, forest, inland wetlands, and shrubland[298]

Diets: Grass and seeds[166]
Scolomys
(spiny mouse)
Anthony, 1924

Two species
  • S. melanops (South American spiny mouse)
  • S. ucayalensis (Ucayali spiny mouse)
Western South America Size range: 8 cm (3 in) long, plus 5 cm (2 in) tail (South American spiny mouse) to 12 cm (5 in) long, plus 8 cm (3 in) tail (Ucayali spiny mouse)[169]

Habitats: Forest[299]

Diets: Seeds and invertebrates[166]
Sigmodon
(cotton rat)

Gray rat

Say & Ord, 1825

Fourteen species
  • S. alleni (Allen's cotton rat)
  • S. alstoni (Alston's cotton rat)
  • S. arizonae (Arizona cotton rat)
  • S. fulviventer (Tawny-bellied cotton rat)
  • S. hirsutus (Southern cotton rat)
  • S. hispidus (Hispid cotton rat)
  • S. inopinatus (Unexpected cotton rat)
  • S. leucotis (White-eared cotton rat, pictured)
  • S. mascotensis (Jaliscan cotton rat)
  • S. ochrognathus (Yellow-nosed cotton rat)
  • S. peruanus (Peruvian cotton rat)
  • S. planifrons (Miahuatlán cotton rat)
  • S. toltecus (Toltec cotton rat)
  • S. zanjonensis (Montane cotton rat)
North America, Central America, and Northern South America Size range: 10 cm (4 in) long, plus 7 cm (3 in) tail (Alston's cotton rat) to 24 cm (9 in) long, plus 16 cm (6 in) tail (Arizona cotton rat)[169]

Habitats: Inland wetlands, grassland, forest, desert, and shrubland[300]

Diets: Omnivorous, including vegetation, insects, invertebrates, and small animals[166]
Sigmodontomys Allen, 1897

One species
  • S. alfari (Alfaro's rice water rat)
Central America and northwestern South America Size: 12–18 cm (5–7 in) long, plus 14–19 cm (6–7 in) tail[169]

Habitats: Inland wetlands and forest[301]

Diet: Seeds and vegetation[301]
Sooretamys Weksler, Percequillo, & Voss, 2006

One species
  • S. angouya (Rat-headed rice rat)
Southeastern South America Size: 12–21 cm (5–8 in) long, plus 16–24 cm (6–9 in) tail[169]

Habitats: Forest and savanna[302]

Diet: Vegetation, seeds, and insects[166]
Tanyuromys Pine, Timm, & Weksler, 2012

One species
  • T. aphrastus (Harris's rice water rat)
Southern Central America and northwestern South America Size: 11–16 cm (4–6 in) long, plus 17–23 cm (7–9 in) tail[169]

Habitats: Forest[303]

Diet: Vegetation, seeds, and insects[166]
Tapecomys

Brown rat

Anderson & Yates, 2000

One species
  • T. primus (Primordial tapecua)
Southern Bolivia and northern Argentina Size: 12–16 cm (5–6 in) long, plus 14–16 cm (6 in) tail[169]

Habitats: Forest[304]

Diet: Vegetation, seeds, and insects[166]
Thalpomys
(cerrado mouse)

Brown mouse

Thomas, 1916

Two species
  • T. cerradensis (Cerrado mouse, pictured)
  • T. lasiotis (Hairy-eared cerrado mouse)
Brazil Size range: 8 cm (3 in) long, plus 5 cm (2 in) tail (hairy-eared cerrado mouse) to 11 cm (4 in) long, plus 7 cm (3 in) tail (Cerrado mouse)[169]

Habitats: Savanna, grassland, and shrubland[305]

Diets: Vegetation, seeds, and insects[166]
Thaptomys

Gray mouse

Thomas, 1916

One species
  • T. nigrita (Blackish grass mouse)
Southeastern South America Size: 8–10 cm (3–4 in) long, plus 3–5 cm (1–2 in) tail[169]

Habitats: Grassland, shrubland, and forest[306]

Diet: Plants and invertebrates[166]
Thomasomys
(Oldfield mouse)

Brown mouse

Coues, 1884

42 species
  • T. andersoni (Anderson's Oldfield mouse)
  • T. apeco (Apeco Oldfield mouse)
  • T. aureus (Golden Oldfield mouse)
  • T. auricularis (Red Andean Oldfield mouse)
  • T. australis (Austral Oldfield mouse)
  • T. baeops (Beady-eyed mouse)
  • T. bombycinus (Silky Oldfield mouse)
  • T. caudivarius (White-tipped Oldfield mouse)
  • T. cinereiventer (Ashy-bellied Oldfield mouse)
  • T. cinereus (Ash-colored Oldfield mouse)
  • T. cinnameus (Cinnamon-colored Oldfield mouse)
  • T. contradictus (Central Andes Oldfield mouse)
  • T. daphne (Daphne's Oldfield mouse)
  • T. dispar (Colombian Oldfield mouse)
  • T. eleusis (Peruvian Oldfield mouse)
  • T. emeritus (Venezuelan Oldfield mouse)
  • T. erro (Wandering Oldfield mouse)
  • T. fumeus (Smoky Oldfield mouse)
  • T. gracilis (Slender Oldfield mouse)
  • T. hudsoni (Hudson's Oldfield mouse)
  • T. hylophilus (Woodland Oldfield mouse)
  • T. incanus (Inca Oldfield mouse)
  • T. ischyrus (Strong-tailed Oldfield mouse)
  • T. kalinowskii (Kalinowski's Oldfield mouse)
  • T. ladewi (Ladew's Oldfield mouse)
  • T. laniger (Soft-furred Oldfield mouse)
  • T. macrotis (Large-eared Oldfield mouse)
  • T. monochromos (Unicolored Oldfield mouse)
  • T. niveipes (Snow-footed Oldfield mouse)
  • T. notatus (Distinguished Oldfield mouse)
  • T. onkiro (Ashaninka Oldfield mouse)
  • T. oreas (Montane Oldfield mouse)
  • T. paramorum (Paramo Oldfield mouse)
  • T. popayanus (Popayán Oldfield mouse)
  • T. praetor (Cajamarca Oldfield mouse)
  • T. pyrrhonotus (Thomas's Oldfield mouse)
  • T. rosalinda (Rosalinda's Oldfield mouse)
  • T. silvestris (Forest Oldfield mouse)
  • T. taczanowskii (Taczanowski's Oldfield mouse, pictured)
  • T. ucucha (Ucucha Oldfield mouse)
  • T. vestitus (Dressy Oldfield mouse)
  • T. vulcani (Pichincha Oldfield mouse)
Northwestern South America Size range: 8 cm (3 in) long, plus 14 cm (6 in) tail (Austral Oldfield mouse) to 24 cm (9 in) long, plus 33 cm (13 in) tail (Apeco Oldfield mouse)[169]

Habitats: Rocky areas, inland wetlands, grassland, forest, and shrubland[307]

Diets: Vegetation, seeds, and insects[166]
Transandinomys
(transandean rice rat)
Weksler, Percequillo, & Voss, 2006

Two species
Central America and northwestern South America Size range: 10 cm (4 in) long, plus 9 cm (4 in) tail (long-whiskered rice rat) to 16 cm (6 in) long, plus 16 cm (6 in) tail (Talamancan rice rat)[169]

Habitats: Forest and inland wetlands[308]

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish[166]
Wiedomys
(red-nosed mouse)

Brown and white mouse

Hershkovitz, 1959

Two species
  • W. cerradensis (Cerrado red-nosed mouse, pictured)
  • W. pyrrhorhinos (Red-nosed mouse)
Eastern Brazil Size range: 10 cm (4 in) long, plus 14 cm (6 in) tail (Cerrado red-nosed mouse) to 12 cm (5 in) long, plus 19 cm (7 in) tail (red-nosed mouse)[169]

Habitats: Savanna, grassland, forest, and shrubland[309]

Diets: Seeds and insects[166]
Wilfredomys

Brown mouse

Avila-Pires, 1960

One species
  • W. oenax (Greater Wilfred's mouse)
Southeastern South America Size: 11–13 cm (4–5 in) long, plus 16–21 cm (6–8 in) tail[169]

Habitats: Forest[310]

Diet: Vegetation, seeds, and insects[166]
Zygodontomys
(cane mouse)
Allen, 1897

Two species
Southern Central America and northern South America Size range: 13 cm (5 in) long, plus 11 cm (4 in) tail (brown cane mouse) to 17 cm (7 in) long, plus 12 cm (5 in) tail (short-tailed cane mouse)[169]

Habitats: Savanna, grassland, forest, and inland wetlands[311]

Diets: Seeds, grass, and fruit[166]
Subfamily Tylomyinae – Reig, 1984 – four genera
Name Authority and species Range Size and ecology
Nyctomys

Brown rat

Saussure, 1860

One species
  • N. sumichrasti (Sumichrast's vesper rat)
Southern Mexico, Central America
Map of range
Size: 12–14 cm (5–6 in) long, plus 14–16 cm (6 in) tail[312]

Habitats: Forest[313]

Diet: Seeds, fruit, and other vegetation[167]
Otonyctomys

Brown rat

Anthony, 1932

One species
Eastern Mexico and northern Central America
Map of range
Size: 9–12 cm (4–5 in) long, plus 6–13 cm (2–5 in) tail[312]

Habitats: Forest[314]

Diet: Vegetation[167]
Ototylomys

Gray rat

Merriam, 1901

One species
  • O. phyllotis (Big-eared climbing rat)
Mexico and Central America Size: 12–17 cm (5–7 in) long, plus 10–18 cm (4–7 in) tail[312]

Habitats: Forest[315]

Diet: Fruit and leaves[167]
Tylomys
(climbing rat)

Gray rat

Peters, 1866

Seven species
  • T. bullaris (Chiapan climbing rat)
  • T. fulviventer (Fulvous-bellied climbing rat)
  • T. mirae (Mira climbing rat)
  • T. nudicaudus (Peters's climbing rat)
  • T. panamensis (Panamanian climbing rat)
  • T. tumbalensis (Tumbala climbing rat)
  • T. watsoni (Watson's climbing rat)
Mexico, Central America, and northwestern South America Size range: 15–29 cm (6–11 in) long, plus 15–32 cm (6–13 in) tail (Watson's climbing rat)[312]

Habitats: Forest[316]

Diets: Seeds, fruit, and other vegetation[167]
Family Muridae

Members of the Muridae family are called murids and include gerbils and many types of mice and rats. They eat a variety of vegetation and invertebrates, though some also eat small vertebrates.[317][318][319][320][321] Muridae comprises 801 extant species, the most of any mammal family, divided into 156 genera. These genera are divided into five subfamilies: Deomyinae, containing 55 species of spiny mice and brush-furred rats; Gerbillinae, containing 102 species of gerbils, jirds, and sand rats; Leimacomyinae, containing a single rat species; Lophiomyinae, containing a single mouse species; and Murinae, containing 642 species of Old World mice and rats. Additionally, thirteen species in Murinae have been driven extinct since 1500.[322]

Subfamily Deomyinae – Thomas, 1888 – four genera
Name Authority and species Range Size and ecology
Acomys
(spiny mouse)

Light brown mouse

I. Geoffroy, 1838

22 species
  • A. airensis (Western Saharan spiny mouse)
  • A. cahirinus (Cairo spiny mouse)
  • A. chudeaui (Chudeau's spiny mouse)
  • A. cilicicus (Asia Minor spiny mouse)
  • A. cineraceus (Gray spiny mouse)
  • A. dimidiatus (Eastern spiny mouse, pictured)
  • A. ignitus (Fiery spiny mouse)
  • A. johannis (Johan's spiny mouse)
  • A. kempi (Kemp's spiny mouse)
  • A. louisae (Louise's spiny mouse)
  • A. minous (Crete spiny mouse)
  • A. mullah (Mullah spiny mouse)
  • A. muzei (Muze spiny mouse)
  • A. nesiotes (Cyprus spiny mouse)
  • A. ngurui (Nguru spiny mouse)
  • A. percivali (Percival's spiny mouse)
  • A. russatus (Golden spiny mouse)
  • A. selousi (Selous's spiny mouse)
  • A. seurati (Seurat's spiny mouse)
  • A. spinosissimus (Southern African spiny mouse)
  • A. subspinosus (Cape spiny mouse)
  • A. wilsoni (Wilson's spiny mouse)
Africa, southern Europe, and southwestern Asia Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (Cape spiny mouse) to 14 cm (6 in) long, plus 12 cm (5 in) tail (Cyprus spiny mouse)[323]

Habitats: Savanna, forest, shrubland, grassland, desert, and rocky areas[324]

Diets: Omnivorous, but primarily grain, grass, and other plant material[317]
Deomys Thomas, 1888

One species
Central Africa Size: 10–15 cm (4–6 in) long, plus 15–20 cm (6–8 in) tail[323]

Habitats: Inland wetlands and forest[325]

Diet: Insects and other invertebrates, as well as vegetable matter[325]
Lophuromys
(brush-furred rat)

Brown mouse

Peters, 1874

31 species
  • L. angolensis (Angolan brush-furred rat)
  • L. ansorgei (Ansorge's brush-furred rat)
  • L. aquilus (Gray brush-furred rat)
  • L. brevicaudus (Short-tailed brush-furred rat)
  • L. brunneus (Thomas's Ethiopian brush-furred rat)
  • L. chercherensis (Mount Chercher brush-furred rat)
  • L. chrysopus (Ethiopian forest brush-furred rat)
  • L. cinereus (Dark-colored brush-furred rat)
  • L. dieterleni (Dieterlen's brush-furred mouse)
  • L. dudui (Dudu's brush-furred rat)
  • L. eisentrauti (Mount Lefo brush-furred mouse)
  • L. flavopunctatus (Yellow-spotted brush-furred rat)
  • L. huttereri (Hutterer's brush-furred mouse)
  • L. kilonzoi (Kilonzo's brush furred rat)
  • L. luteogaster (Yellow-bellied brush-furred rat)
  • L. machangui (Machandu's brush furred rat)
  • L. makundii (Makundi's brush-furred rat)
  • L. medicaudatus (Medium-tailed brush-furred rat)
  • L. melanonyx (Black-clawed brush-furred rat)
  • L. menageshae (North Western Rift brush-furred rat)
  • L. nudicaudus (Fire-bellied brush-furred rat)
  • L. pseudosikapusi (Sheko Forest brush-furred rat)
  • L. rahmi (Rahm's brush-furred rat)
  • L. roseveari (Mount Cameroon brush-furred rat)
  • L. sabunii (Sabuni's brush-furred rat)
  • L. sikapusi (Rusty-bellied brush-furred rat, pictured)
  • L. simensis (Simien brush-furred rat)
  • L. stanleyi (Stanley's brush-furred rat)
  • L. verhageni (Verhagen's brush-furred rat)
  • L. woosnami (Woosnam's brush-furred rat)
  • L. zena (Zena's brush-furred rat)
Western, central, and eastern Africa Size range: 8 cm (3 in) long, plus 5 cm (2 in) tail (Dudu's brush-furred rat) to 18 cm (7 in) long, plus 10 cm (4 in) tail (black-clawed brush-furred rat)[323]

Habitats: Rocky areas, grassland, forest, and shrubland[326]

Diets: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[317]
Uranomys

Brown mouse

Dollman, 1909

One species
  • U. ruddi (Rudd's mouse)
Western, central, and eastern Africa Size: 9–12 cm (4–5 in) long, plus 6–8 cm (2–3 in) tail[323]

Habitats: Savanna and shrubland[327]

Diet: Insects[317]
Subfamily GerbillinaeGray, 1825 – fourteen genera
Name Authority and species Range Size and ecology
Ammodillus Thomas, 1904

One species
Ethiopia and Somalia Size: 8–11 cm (3–4 in) long, plus 12–15 cm (5–6 in) tail[328]

Habitats: Grassland[329]

Diet: Plant material and invertebrates[318]
Brachiones

Drawing of brown gerbils

Thomas, 1925

One species
Northern China Size: 6–11 cm (2–4 in) long, plus 5–8 cm (2–3 in) tail[328]

Habitats: Desert[330]

Diet: Plant material and invertebrates[318]
Desmodilliscus Wettstein, 1916

One species
Western and northern Africa
Map of range
Size: 5–8 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[328]

Habitats: Savanna[331]

Diet: Grain[318]
Desmodillus

Brown gerbil

Thomas & Schwann, 1904

One species
Southern Africa Size: 8–13 cm (3–5 in) long, plus 7–10 cm (3–4 in) tail[328]

Habitats: Desert[332]

Diet: Seeds, grain, and insects[318]
Gerbilliscus
(large naked-soled gerbil)

Brown gerbil

Thomas, 1897

Sixteen species
  • G. afra (Cape gerbil)
  • G. boehmi (Boehm's gerbil)
  • G. brantsii (Highveld gerbil)
  • G. gambiana (Sahelo-Sudanian gerbil)
  • G. guineae (Guinean gerbil)
  • G. inclusus (Gorongoza gerbil)
  • G. kempi (Kemp's gerbil)
  • G. leucogaster (Bushveld gerbil, pictured)
  • G. nigricaudus (Black-tailed gerbil)
  • G. paeba (Hairy-footed gerbil)
  • G. phillipsi (Phillips's gerbil)
  • G. robustus (Fringe-tailed gerbil)
  • G. setzeri (Namib brush-tailed gerbil)
  • G. tytonis (Dune hairy-footed gerbil)
  • G. validus (Savanna gerbil)
  • G. vallinus (Bushy-tailed hairy-footed gerbil)
Sub-Saharan Africa Size range: 8 cm (3 in) long, plus 12 cm (5 in) tail (bushveld gerbil) to 20 cm (8 in) long, plus 21 cm (8 in) tail (black-tailed gerbil)[328]

Habitats: Desert, forest, savanna, grassland, shrubland, and rocky areas[333]

Diets: Roots, bulbs, seeds, vegetation, and insects; Seeds[318]
Gerbillus
(northern pygmy gerbil)

Brown and white gerbil

Desmarest, 1804

49 species
  • G. acticola (Berbera gerbil)
  • G. agag (Agag gerbil)
  • G. amoenus (Pleasant gerbil)
  • G. andersoni (Anderson's gerbil)
  • G. aquilus (Swarthy gerbil)
  • G. bottai (Botta's gerbil)
  • G. brockmani (Brockman's gerbil)
  • G. burtoni (Burton's gerbil)
  • G. campestris (North African gerbil)
  • G. cheesmani (Cheesman's gerbil, pictured)
  • G. dasyurus (Wagner's gerbil)
  • G. dunni (Dunn's gerbil)
  • G. famulus (Black-tufted gerbil)
  • G. floweri (Flower's gerbil)
  • G. garamantis (Algerian gerbil)
  • G. gerbillus (Lesser Egyptian gerbil)
  • G. gleadowi (Indian hairy-footed gerbil)
  • G. grobbeni (Grobben's gerbil)
  • G. harwoodi (Harwood's gerbil)
  • G. henleyi (Pygmy gerbil)
  • G. hesperinus (Western gerbil)
  • G. hoogstraali (Hoogstraal's gerbil)
  • G. jamesi (James's gerbil)
  • G. latastei (Lataste's gerbil)
  • G. lowei (Lowe's gerbil)
  • G. mackilligini (Mackilligin's gerbil)
  • G. maghrebi (Greater short-tailed gerbil)
  • G. mauritaniae (Mauritanian gerbil)
  • G. mesopotamiae (Harrison's gerbil)
  • G. muriculus (Darfur gerbil)
  • G. nancillus (Sudan gerbil)
  • G. nanus (Balochistan gerbil)
  • G. nigeriae (Nigerian gerbil)
  • G. occiduus (Occidental gerbil)
  • G. perpallidus (Pale gerbil)
  • G. poecilops (Large Aden gerbil)
  • G. principulus (Principal gerbil)
  • G. pulvinatus (Cushioned gerbil)
  • G. pusillus (Least gerbil)
  • G. pyramidum (Greater Egyptian gerbil)
  • G. rosalinda (Rosalinda gerbil)
  • G. rupicola (Rupicolous gerbil)
  • G. simoni (Lesser short-tailed gerbil)
  • G. somalicus (Somalian gerbil)
  • G. stigmonyx (Khartoum gerbil)
  • G. syrticus (Sand gerbil)
  • G. tarabuli (Tarabul's gerbil)
  • G. vivax (Vivacious gerbil)
  • G. watersi (Waters's gerbil)
Northern, central, and eastern Africa and western and southern Asia Size range: 5 cm (2 in) long, plus 8 cm (3 in) tail (least gerbil) to 22 cm (9 in) long, plus 11 cm (4 in) tail (large Aden gerbil)[328]

Habitats: Desert, savanna, forest, grassland, shrubland, coastal marine, and rocky areas[334]

Diets: Seeds, roots, nuts, grass, and insects[318]
Meriones
(jird)

Brown and white gerbil

Illiger, 1811

Seventeen species
  • M. arimalius (Arabian jird)
  • M. chengi (Cheng's jird)
  • M. crassus (Sundevall's jird)
  • M. dahli (Dahl's jird)
  • M. grandis (Moroccan jird)
  • M. hurrianae (Indian desert jird)
  • M. libycus (Libyan jird)
  • M. meridianus (Midday jird)
  • M. persicus (Persian jird)
  • M. rex (King jird)
  • M. sacramenti (Buxton's jird)
  • M. shawi (Shaw's jird)
  • M. tamariscinus (Tamarisk jird)
  • M. tristrami (Tristram's jird)
  • M. unguiculatus (Mongolian gerbil, pictured)
  • M. vinogradovi (Vinogradov's jird)
  • M. zarudnyi (Zarudny's jird)
Northern Africa and western, central, and southern Asia Size range: 9 cm (4 in) long, plus 8 cm (3 in) tail (Mongolian gerbil) to 20 cm (8 in) long, plus 18 cm (7 in) tail (Moroccan jird)[328]

Habitats: Desert, savanna, grassland, inland wetlands, shrubland, and rocky areas[335]

Diets: Vegetation, roots, bulbs, seeds, grain, fruit, and insects[318]
Microdillus Thomas, 1910

One species
  • M. peeli (Somali pygmy gerbil)
Somalia Size: 6–8 cm (2–3 in) long, plus 5–7 cm (2–3 in) tail[328]

Habitats: Grassland and shrubland[336]

Diet: Seeds, roots, nuts, grass, and insects[318]
Pachyuromys

Brown gerbils

Lataste, 1880

One species
Northern Africa Size: 9–12 cm (4–5 in) long, plus 5–7 cm (2–3 in) tail[328]

Habitats: Shrubland and desert[337]

Diet: Insects[318]
Psammomys
(sand rat)

Brown gerbil

Cretzschmar, 1828

Two species
  • P. obesus (Fat sand rat, pictured)
  • P. vexillaris (Thin sand rat)
Northern Africa and Middle East Size range: 11 cm (4 in) long, plus 8 cm (3 in) tail (thin sand rat) to 19 cm (7 in) long, plus 14 cm (6 in) tail (fat sand rat)[328]

Habitats: Inland wetlands, desert, and shrubland[338]

Diets: Succulents and other plants[318]
Rhombomys

Brown gerbil

Wagner, 1841

One species
Central Asia Size: 15–19 cm (6–7 in) long, plus 13–16 cm (5–6 in) tail[328]

Habitats: Desert, grassland, and shrubland[339]

Diet: Variety of desert plants[318]
Sekeetamys

Brown gerbil

Ellerman, 1947

One species
Egypt and Middle East Size: 9–13 cm (4–5 in) long, plus 11–17 cm (4–7 in) tail[328]

Habitats: Rocky areas and desert[340]

Diet: Plants and insects[340]
Tatera

Brown and white gerbil

Lataste, 1882

One species
Southern and western Asia Size: 14–21 cm (6–8 in) long, plus 15–20 cm (6–8 in) tail[328]

Habitats: Forest, desert, grassland, and shrubland[341]

Diet: Roots, bulbs, seeds, vegetation, insects, eggs, and birds[318]
Taterillus
(tateril)

Brown and white gerbil

Thomas, 1910

Nine species
  • T. arenarius (Robbins's tateril)
  • T. congicus (Congo gerbil)
  • T. emini (Emin's gerbil, pictured)
  • T. gracilis (Gracile tateril)
  • T. harringtoni (Harrington's gerbil)
  • T. lacustris (Lake Chad gerbil)
  • T. petteri (Petter's gerbil)
  • T. pygargus (Senegal gerbil)
  • T. tranieri (Tranier's tateril)
Eastern, central, and western Africa Size range: 9 cm (4 in) long, plus 13 cm (5 in) tail (gracile tateril) to 15 cm (6 in) long, plus 19 cm (7 in) tail (Congo gerbil)[328]

Habitats: Grassland, forest, savanna, and shrubland[342]

Diets: Grain and insects[318]
Subfamily LeimacomyinaeMusser & Carleton, 2005 – one genus
Name Authority and species Range Size and ecology
Leimacomys

Light brown mouse

Matschie, 1893

One species
Togo
Map of range
Size: About 12 cm (5 in) long, plus about 4 cm (2 in) tail[343]

Habitats: Unknown[344]

Diet: Vegetation and insects[319]
Subfamily LophiomyinaeMilne-Edwards, 1867 – one genus
Name Authority and species Range Size and ecology
Lophiomys

Light brown mouse

Milne-Edwards, 1867

One species
Eastern Africa Size: 25–30 cm (10–12 in) long, plus 14–20 cm (6–8 in) tail[343]

Habitats: Forest, savanna, and shrubland[345]

Diet: Leaves and shoots[320]
Subfamily MurinaeIlliger, 1811 – 136 genera
Name Authority and species Range Size and ecology
Abditomys Musser, 1982

One species
Philippines
Map of range
Size: 23–27 cm (9–11 in) long, plus 22–24 cm (9 in) tail[322]

Habitats: Forest[346]

Diet: Vegetation[321]
Abeomelomys Menzies, 1990

One species
  • A. sevia (Highland brush mouse)
Papua New Guinea Size: 11–14 cm (4–6 in) long, plus 14–20 cm (6–8 in) tail[322]

Habitats: Grassland and forest[347]

Diet: Plant material and invertebrates[321]
Aethomys
(rock rat)

Brown rat

Thomas, 1915

Nine species
  • A. bocagei (Bocage's rock rat)
  • A. chrysophilus (Red rock rat)
  • A. hindei (Hinde's rock rat)
  • A. ineptus (Tete veld aethomys, pictured)
  • A. kaiseri (Kaiser's rock rat)
  • A. nyikae (Nyika rock rat)
  • A. silindensis (Selinda veld rat)
  • A. stannarius (Tinfields rock rat)
  • A. thomasi (Thomas's rock rat)
Central, eastern, and southern Africa Size range: 12 cm (5 in) long, plus 12 cm (5 in) tail (Hinde's rock rat) to 20 cm (8 in) long, plus 18 cm (7 in) tail (Bocage's rock rat)[322]

Habitats: Savanna, grassland, forest, rocky areas, and shrubland[348]

Diets: Grain, seeds, roots, nuts, and fruit[321]
Anisomys Thomas, 1904

One species
  • A. imitator (Squirrel-toothed rat)
New Guinea Size: 21–28 cm (8–11 in) long, plus 27–36 cm (11–14 in) tail[322]

Habitats: Forest[349]

Diet: Nuts and other vegetation[321]
Anonymomys Musser, 1981

One species
Philippines
Map of range
Size: 14–15 cm (6 in) long, plus 19–22 cm (7–9 in) tail[322]

Habitats: Forest[350]

Diet: Plant material and invertebrates[321]
Apodemus
(field mouse)

Brown and white mouse

Kaup, 1829

20 species
  • A. agrarius (Striped field mouse)
  • A. alpicola (Alpine field mouse)
  • A. argenteus (Small Japanese field mouse, pictured)
  • A. chevrieri (Chevrier's field mouse)
  • A. draco (South China field mouse)
  • A. epimelas (Western broad-toothed field mouse)
  • A. flavicollis (Yellow-necked mouse)
  • A. gurkha (Himalayan field mouse)
  • A. hyrcanicus (Caucasus field mouse)
  • A. latronum (Sichuan field mouse)
  • A. mystacinus (Eastern broad-toothed field mouse)
  • A. pallipes (Ward's field mouse)
  • A. peninsulae (Korean field mouse)
  • A. ponticus (Black Sea field mouse)
  • A. rusiges (Kashmir field mouse)
  • A. semotus (Taiwan field mouse)
  • A. speciosus (Large Japanese field mouse)
  • A. sylvaticus (Wood mouse)
  • A. uralensis (Ural field mouse)
  • A. witherbyi (Steppe field mouse)
Europe, Asia, and northern Africa Size range: 6 cm (2 in) long, plus 7 cm (3 in) tail (small Japanese field mouse) to 15 cm (6 in) long, plus 15 cm (6 in) tail (western broad-toothed field mouse)[322]

Habitats: Coastal marine, grassland, inland wetlands, forest, rocky areas, and shrubland[351]

Diets: Roots, grain, seeds, berries, nuts, and insects[321]
Apomys
(earthworm mouse)

Dead brown mouse

Mearns, 1905

Nineteen species
  • A. abrae (Luzon Cordillera forest mouse)
  • A. aurorae (Luzon Aurora forest mouse)
  • A. banahao (Mount Banahaw forest mouse)
  • A. brownorum (Mount Tapulao forest mouse)
  • A. camiguinensis (Camiguin forest mouse)
  • A. datae (Luzon montane forest mouse)
  • A. gracilirostris (Large Mindoro forest mouse)
  • A. hylocetes (Mount Apo forest mouse)
  • A. insignis (Mindanao montane forest mouse)
  • A. iridensis (Mount Irid forest mouse)
  • A. littoralis (Mindanao lowland forest mouse)
  • A. lubangensis (Lubang forest mouse)
  • A. magnus (Luzon giant forest mouse)
  • A. microdon (Small Luzon forest mouse)
  • A. minganensis (Mount Mingan forest mouse)
  • A. musculus (Least forest mouse)
  • A. sacobianus (Long-nosed Luzon forest mouse, pictured)
  • A. sierrae (Sierra Madre forest mouse)
  • A. zambalensis (Luzon Zambales forest mouse)
Philippines Size range: 7 cm (3 in) long, plus 9 cm (4 in) tail (least forest mouse) to 18 cm (7 in) long, plus 16 cm (6 in) tail (Sierra Madre forest mouse)[322]

Habitats: Forest, shrubland, and grassland[352]

Diets: Omnivorous[321]
Archboldomys
(shrew-mouse)
Musser, 1982

Two species
  • A. luzonensis (Mount Isarog shrew-mouse)
  • A. maximus (Large Cordillera shrew-mouse)
Philippines Size range: 10–12 cm (4–5 in) long, plus 9–11 cm (4 in) tail (large Cordillera shrew-mouse)[322]

Habitats: Forest[353]

Diets: Worms and soft-bodied invertebrates[321]
Arvicanthis
(unstriped grass rat)

Brown rat

Lesson, 1842

Seven species
  • A. abyssinicus (Abyssinian grass rat)
  • A. ansorgei (Sudanian grass rat)
  • A. blicki (Blick's grass rat)
  • A. nairobae (Nairobi grass rat)
  • A. neumanni (Neumann's grass rat)
  • A. niloticus (African grass rat, pictured)
  • A. rufinus (Guinean grass rat)
Africa Size range: 11 cm (4 in) long, plus 12 cm (5 in) tail (Guinean grass rat) to 19 cm (7 in) long, plus 16 cm (6 in) tail (African grass rat)[322]

Habitats: Savanna, shrubland, and grassland[354]

Diets: Seeds, leaves, grass, and grain[321]
Baiyankamys
(mountain water rat)
Hinton, 1943

Two species
  • B. habbema (Mountain water rat)
  • B. shawmayeri (Shaw Mayer's water rat)
New Guinea
Map of range
Size range: 13–16 cm (5–6 in) long, plus 15–19 cm (6–7 in) tail (mountain water rat)[322]

Habitats: Forest, inland wetlands, grassland, and shrubland[355]

Diets: Fish and aquatic insects, as well as spiders, crustaceans, mussels, frogs, turtles, birds, and bats[321]
Bandicota
(bandicoot rat)

Brown rat

Gray, 1873

Three species
Southern and southeastern Asia Size range: 14 cm (6 in) long, plus 7 cm (3 in) tail (Savile's bandicoot rat) to 35 cm (14 in) long, plus 28 cm (11 in) tail (greater bandicoot rat)[322]

Habitats: Forest, inland wetlands, shrubland, and grassland[356]

Diets: Omnivorous[321]
Batomys
(hairy-tailed rat)

Gray rat

Thomas, 1895

Six species
  • B. dentatus (Large-toothed hairy-tailed rat)
  • B. granti (Luzon Cordillera hairy-tailed rat)
  • B. hamiguitan (Hamiguitan hairy-tailed rat)
  • B. russatus (Dinagat hairy-tailed rat)
  • B. salomonseni (Mindanao hairy-tailed rat)
  • B. uragon (Mount Isarog hairy-tailed rat)
Philippines
Map of range
Size range: 13 cm (5 in) long, plus 13 cm (5 in) tail (Mindanao hairy-tailed rat) to 21 cm (8 in) long, plus 18 cm (7 in) tail (Mount Isarog hairy-tailed rat)[322]

Habitats: Forest[357]

Diets: Leaves, seeds, and fruit[321]
Berylmys
(white-toothed rat)

Drawing of brown rat

Ellerman, 1947

Five species
  • B. berdmorei (Small white-toothed rat)
  • B. bowersi (Bower's white-toothed rat, pictured)
  • B. latouchei (West Chinese white-toothed rat)
  • B. mackenziei (Kenneth's white-toothed rat)
  • B. manipulus (Manipur white-toothed rat)
Southeastern Asia and eastern China Size range: 13 cm (5 in) long, plus 14 cm (6 in) tail (Manipur white-toothed rat) to 30 cm (12 in) long, plus 31 cm (12 in) tail (Bower's white-toothed rat)[322]

Habitats: Forest, grassland, and shrubland[358]

Diets: Leaves, grass, seeds, fruit, insects, molluscs, and worms[321]
Brassomys Musser & Lunde, 2009

One species
  • B. albidens (White-toothed brush mouse)
Western New Guinea
Map of range
Size: 11–13 cm (4–5 in) long, plus 14–17 cm (6–7 in) tail[322]

Habitats: Grassland and shrubland[359]

Diet: Leaves and other vegetation[321]
Bullimus
(large forest rat)

Gray rat

Mearns, 1905

Three species
  • B. bagobus (Bagobo rat, pictured)
  • B. gamay (Camiguin forest rat)
  • B. luzonicus (Large Luzon forest rat)
Philippines Size range: 22 cm (9 in) long, plus 14 cm (6 in) tail (Camiguin forest rat) to 27 cm (11 in) long, plus 20 cm (8 in) tail (Bagobo rat)[322]

Habitats: Forest and shrubland[360]

Diets: Plant material and invertebrates[321]
Bunomys
(hill rat)

Dead brown rat

Thomas, 1910

Seven species
  • B. andrewsi (Andrew's hill rat)
  • B. chrysocomus (Yellow-haired hill rat, pictured)
  • B. coelestis (Heavenly hill rat)
  • B. karokophilus (Karoko hill rat)
  • B. penitus (Inland hill rat)
  • B. prolatus (Long-headed hill rat)
  • B. torajae (Tana Toraja hill rat)
Indonesia Size range: 9 cm (4 in) long, plus 9 cm (4 in) tail (yellow-haired hill rat) to 24 cm (9 in) long, plus 19 cm (7 in) tail (inland hill rat)[322]

Habitats: Forest and shrubland[361]

Diets: Fruit, insects, snails, and worms[321]
Carpomys
(Luzon tree rat)

Drawing of brown and white rat

Thomas, 1895

Two species
Philippines Size range: 16 cm (6 in) long, plus 15 cm (6 in) tail (white-bellied Luzon tree rat) to 19 cm (7 in) long, plus 19 cm (7 in) tail (short-footed Luzon tree rat)[322]

Habitats: Forest[362]

Diets: Plant material and invertebrates[321]
Chiromyscus Thomas, 1925

One species
  • C. chiropus (Fea's tree rat)
Southeastern Asia
Map of range
Size: 13–16 cm (5–6 in) long, plus 20–23 cm (8–9 in) tail[322]

Habitats: Shrubland and forest[363]

Diet: Plant material and invertebrates[321]
Chiropodomys
(pencil-tailed tree mouse)
Peters, 1869

Six species
  • C. calamianensis (Palawan pencil-tailed tree mouse)
  • C. gliroides (Indomalayan pencil-tailed tree mouse)
  • C. karlkoopmani (Koopman's pencil-tailed tree mouse)
  • C. major (Large pencil-tailed tree mouse)
  • C. muroides (Gray-bellied pencil-tailed tree mouse)
  • C. pusillus (Small pencil-tailed tree mouse)
Southeastern Asia Size range: 6 cm (2 in) long, plus 8 cm (3 in) tail (gray-bellied pencil-tailed tree mouse) to 13 cm (5 in) long, plus 18 cm (7 in) tail (Palawan pencil-tailed tree mouse)[322]

Habitats: Forest[364]

Diets: Vegetation[321]
Chiruromys
(tree mouse)
Thomas, 1888

Three species
  • C. forbesi (Greater tree mouse)
  • C. lamia (Broad-headed tree mouse)
  • C. vates (Lesser tree mouse)
Papua New Guinea Size range: 8 cm (3 in) long, plus 12 cm (5 in) tail (lesser tree mouse) to 17 cm (7 in) long, plus 23 cm (9 in) tail (greater tree mouse)[322]

Habitats: Forest[365]

Diets: Leaves and grass and bamboo shoots[321]
Chrotomys
(striped shrew-rat)

Drawing of brown, yellow, and white rat

Thomas, 1895

Five species
  • C. gonzalesi (Isarog striped shrew-rat)
  • C. mindorensis (Mindoro striped rat, pictured)
  • C. sibuyanensis (Sibuyan striped shrew-rat)
  • C. silaceus (Blazed Luzon shrew-rat)
  • C. whiteheadi (Luzon striped rat)
Philippines Size range: 13 cm (5 in) long, plus 9 cm (4 in) tail (blazed Luzon shrew-rat) to 19 cm (7 in) long, plus 11 cm (4 in) tail (Isarog striped shrew-rat)[322]

Habitats: Forest[366]

Diets: Sweet potatoes, grass, and worms[321]
Coccymys
(brush mouse)
Menzies, 1990

Three species
  • C. kirrhos (Tawny brush mouse)
  • C. ruemmleri (Rümmler's brush mouse)
  • C. shawmayeri (Central Cordillera brush mouse)
Papua New Guinea and New Guinea Size range: 8 cm (3 in) long, plus 13 cm (5 in) tail (Central Cordillera brush mouse) to 12 cm (5 in) long, plus 18 cm (7 in) tail (Rümmler's brush mouse)[322]

Habitats: Forest and grassland[367]

Diets: Leaves and other vegetation[321]
Colomys Thomas & Wroughton, 1907

One species
Central Africa Size: 10–16 cm (4–6 in) long, plus 14–19 cm (6–7 in) tail[322]

Habitats: Savanna, forest, grassland, and inland wetlands[368]

Diet: Worms, slugs, crustaceans, and aquatic insects, as well as small vertebrates and some vegetation[321]
Conilurus
(rabbit rat)

Drawing of brown and white rat

Ogilby, 1838

Three species
  • C. albipes (White-footed rabbit rat)
  • C. capricornensis (Capricorn rabbit rat)
  • C. penicillatus (Brush-tailed rabbit rat)
Southeastern Australia, Northern Australia and Papua New Guinea, and Northeastern Australia Size range: 15–20 cm (6–8 in) long, plus 17–21 cm (7–8 in) tail (brush-tailed rabbit rat)[322]

Habitats: Savanna and forest[369]

Diets: Plant material and invertebrates[321]
Crateromys
(bushy-tailed cloud rat)

Brown rat

Thomas, 1895

Four species
Philippines Size range: 25 cm (10 in) long, plus 21 cm (8 in) tail (Ilin Island cloudrunner) to 37 cm (15 in) long, plus 39 cm (15 in) tail (giant bushy-tailed cloud rat)[322]

Habitats: Forest[370]

Diets: Fruit and pine tree sprouts, buds, and bark[321]
Cremnomys
(Cutch rat)

Wroughton, 1912

Two species
  • C. cutchicus (Cutch rat, pictured)
  • C. elvira (Elvira rat)
India and Southern India Size range: 10 cm (4 in) long, plus 12 cm (5 in) tail (Cutch rat) to 15 cm (6 in) long, plus 20 cm (8 in) tail (Elvira rat)[322]

Habitats: Grassland, desert, forest, rocky areas, and shrubland[371]

Diets: Plant material and invertebrates[321]
Crossomys Thomas, 1907

One species
New Guinea
Map of range
Size: 17–23 cm (7–9 in) long, plus 21–27 cm (8–11 in) tail[322]

Habitats: Forest and inland wetlands[372]

Diet: Tadpoles, insects, mollusks, and small aquatic vertebrates[321]
Crunomys
(shrew-rat)

Drawing of brown and white rat

Thomas, 1897

Four species
  • C. celebensis (Celebes shrew-rat)
  • C. fallax (Northern Luzon shrew-rat, pictured)
  • C. melanius (Mindanao shrew-rat)
  • C. suncoides (Katanglad shrew-mouse)
Philippines and Indonesia Size range: 9–14 cm (4–6 in) long, plus 6–10 cm (2–4 in) tail (Mindanao shrew-rat)[322]

Habitats: Forest[373]

Diets: Plant material and invertebrates[321]
Dacnomys Thomas, 1916

One species
  • D. millardi (Millard's rat)
Southeastern Asia
Map of range
Size: 21–27 cm (8–11 in) long, plus 29–33 cm (11–13 in) tail[322]

Habitats: Forest[374]

Diet: Plant material and invertebrates[321]
Dasymys
(marsh rat)
Peters, 1875

Nine species
  • D. alleni (Glover Allen's dasymys)
  • D. cabrali (Crawford-Cabral's shaggy rat)
  • D. foxi (Fox's shaggy rat)
  • D. incomtus (African marsh rat)
  • D. montanus (Montane shaggy rat)
  • D. nudipes (Angolan marsh rat)
  • D. rufulus (West African shaggy rat)
  • D. rwandae (Rwandan shaggy rat)
  • D. sua (Tanzanian shaggy rat)
Sub-Saharan Africa
Map of range
Size range: 11 cm (4 in) long, plus 10 cm (4 in) tail (Glover Allen's dasymys) to 20 cm (8 in) long, plus 16 cm (6 in) tail (African marsh rat)[322]

Habitats: Savanna, grassland, inland wetlands, forest, and shrubland[375]

Diets: Aquatic plants, as well as insects[321]
Dephomys
(Ivory Coast rat)
Thomas, 1926

Two species
  • D. defua (Defua rat)
  • D. eburneae (Ivory Coast rat)
Western Africa Size range: 11 cm (4 in) long, plus 18 cm (7 in) tail (Defua rat) to 15 cm (6 in) long, plus 20 cm (8 in) tail (Ivory Coast rat)[322]

Habitats: Forest[376]

Diets: Fruit and insects[321]
Desmomys
(scrub rat)
Thomas, 1910

Two species
  • D. harringtoni (Harrington's rat)
  • D. yaldeni (Yalden's rat)
Ethiopia Size range: 11 cm (4 in) long, plus 14 cm (6 in) tail (Yalden's rat) to 15 cm (6 in) long, plus 14 cm (6 in) tail (Harrington's rat)[322]

Habitats: Forest and shrubland[377]

Diets: Plant material and invertebrates[321]
Diomys Thomas, 1917

One species
  • D. crumpi (Crump's mouse)
Scattered southern Asia
Map of range
Size: 10–14 cm (4–6 in) long, plus 10–13 cm (4–5 in) tail[322]

Habitats: Forest[378]

Diet: Plant material and invertebrates[321]
Diplothrix

Brown rat

Thomas, 1916

One species
  • D. legata (Ryukyu long-tailed giant rat)
Southern islands of Japan Size: About 23 cm (9 in)[322]

Habitats: Forest[379]

Diet: Plant material and invertebrates[321]
Echiothrix
(echiothrix)

Dead brown rat

Gray, 1867

Two species
  • E. centrosa (Central Sulawesi echiothrix, pictured)
  • E. leucura (Northern Sulawesi echiothrix)
Indonesia Size range: 18 cm (7 in) long, plus 23 cm (9 in) tail (central Sulawesi echiothrix) to 23 cm (9 in) long, plus 27 cm (11 in) tail (northern Sulawesi echiothrix)[322]

Habitats: Forest[380]

Diets: Worms[321]
Eropeplus Miller & Hollister, 1921

One species
  • E. canus (Sulawesi soft-furred rat)
Indonesia Size: 23–25 cm (9–10 in) long, plus 27–30 cm (11–12 in) tail[322]

Habitats: Forest[381]

Diet: Plant material and invertebrates[321]
Frateromys Sody, 1941

One species
  • F. fratrorum (Northeastern peninsula hill rat)
Indonesia Size: 15–19 cm (6–7 in) long, plus 15–20 cm (6–8 in) tail[322]

Habitats: Forest[382]

Diet: Plant material and invertebrates[321]
Golunda

Brown rat

Gray, 1837

One species
  • G. ellioti (Indian bush rat)
Southern Asia Size: 9–17 cm (4–7 in) long, plus 7–14 cm (3–6 in) tail[322]

Habitats: Shrubland, forest, and grassland[383]

Diet: Roots and grass, as well as other vegetation[321]
Gracilimus

Brown rat

Rowe, Achmadi, & Esselstyn, 2016

One species
  • G. radix (Sulawesi root rat)
Indonesia Size: 10–13 cm (4–5 in) long, plus 16–17 cm (6–7 in) tail[322]

Habitats: Forest[384]

Diet: Plant material and invertebrates[321]
Grammomys
(thicket rat)

Gray rat

Thomas, 1915

Eleven species
  • G. aridulus (Arid thicket rat)
  • G. buntingi (Bunting's thicket rat)
  • G. caniceps (Gray-headed thicket rat)
  • G. cometes (Mozambique thicket rat)
  • G. dolichurus (Woodland thicket rat, pictured)
  • G. dryas (Forest thicket rat)
  • G. gigas (Giant thicket rat)
  • G. ibeanus (Ruwenzori thicket rat)
  • G. kuru (Eastern rainforest grammomys)
  • G. macmillani (Macmillan's thicket rat)
  • G. minnae (Ethiopian thicket rat)
Sub-Saharan Africa Size range: 8 cm (3 in) long, plus 14 cm (6 in) tail (gray-headed thicket rat) to 15 cm (6 in) long, plus 18 cm (7 in) tail (eastern rainforest grammomys)[322]

Habitats: Savanna, grassland, inland wetlands, forest, and shrubland[385]

Diets: Stems, fruit, nuts, flowers, and other vegetation, as well as insects[321]
Hadromys
(bush rat)

Drawing of brown and white rat

Thomas, 1911

Two species
  • H. humei (Manipur bush rat, pictured)
  • H. yunnanensis (Yunnan bush rat)
Eastern India and Southern China Size range: 9 cm (4 in) long, plus 12 cm (5 in) tail (Manipur bush rat) to 14 cm (6 in) long, plus 14 cm (6 in) tail (Manipur bush rat)[322]

Habitats: Forest[386]

Diets: Grass[321]
Haeromys
(ranee mouse)
Thomas, 1911

Three species
  • H. margarettae (Ranee mouse)
  • H. minahassae (Minahassa ranee mouse)
  • H. pusillus (Lesser ranee mouse)
Southeastern Asia Size range: 7 cm (3 in) long, plus 12 cm (5 in) tail (lesser ranee mouse) to 8 cm (3 in) long, plus 14 cm (6 in) tail (Minahassa ranee mouse)[322]

Habitats: Forest[387]

Diets: Seeds[321]
Halmaheramys
(Halmahera spiny rat)
Fabre, Pagès, Musser, Fitriana, Semiadi, & Helgen, 2013

Two species
  • H. bokimekot (Spiny Boki Mekot rat)
  • H. wallacei (Wallace's large spiny rat)
Indonesia Size range: 14 cm (6 in) long, plus 11 cm (4 in) tail (spiny Boki Mekot rat) to 24 cm (9 in) long, plus 21 cm (8 in) tail (Wallace's large spiny rat)[322]

Habitats: Forest[388]

Diets: Plant material and invertebrates[321]
Hapalomys
(marmoset rat)

Brown rat

Blyth, 1859

Three species
  • H. delacouri (Delacour's marmoset rat, pictured)
  • H. longicaudatus (Marmoset rat)
  • H. suntsovi (Suntsov's marmoset rat)
Southeastern Asia Size range: 10 cm (4 in) long, plus 13 cm (5 in) tail (Delacour's marmoset rat) to 17 cm (7 in) long, plus 20 cm (8 in) tail (marmoset rat)[322]

Habitats: Forest[389]

Diets: Bamboo shoots, flowers, and fruit[321]
Heimyscus Misonne, 1969

One species
  • H. fumosus (African smoky mouse)
West-central Africa Size: 8–10 cm (3–4 in) long, plus 8–12 cm (3–5 in) tail[322]

Habitats: Forest[390]

Diet: Plant material and invertebrates[321]
Hybomys
(striped mouse)
Thomas, 1910

Six species
  • H. badius (Eisentraut's striped mouse)
  • H. basilii (Father Basilio's striped mouse)
  • H. lunaris (Moon striped mouse)
  • H. planifrons (Miller's striped mouse)
  • H. trivirgatus (Temminck's striped mouse)
  • H. univittatus (Peters's striped mouse)
Western Uganda and eastern Democratic Republic of the Congo, Central Africa, Western Africa, and Cameroon Size range: 10 cm (4 in) long, plus 10 cm (4 in) tail (Eisentraut's striped mouse) to 16 cm (6 in) long, plus 14 cm (6 in) tail (Father Basilio's striped mouse)[322]

Habitats: Forest[391]

Diets: Fruit, roots, and insects[321]
Hydromys
(water rat)

Brown rat

Geoffroy, 1804

Four species
  • H. chrysogaster (Rakali, pictured)
  • H. hussoni (Western water rat)
  • H. neobritannicus (New Britain water rat)
  • H. ziegleri (Ziegler's water rat)
Australia and southeastern Asia Size range: 12 cm (5 in) long, plus 10 cm (4 in) tail (western water rat) to 39 cm (15 in) long, plus 32 cm (13 in) tail (rakali)[322]

Habitats: Coastal marine, forest, inland wetlands, and neritic marine[392]

Diets: Fish and aquatic insects, as well as spiders, crustaceans, mussels, frogs, turtles, birds, and bats[321]
Hylomyscus
(wood mouse)
Thomas, 1926

Sixteen species
  • H. aeta (Beaded wood mouse)
  • H. alleni (Allen's wood mouse)
  • H. anselli (Ansell's wood mouse)
  • H. arcimontensis (Arc Mountain wood mouse)
  • H. baeri (Baer's wood mouse)
  • H. carillus (Angolan wood mouse)
  • H. denniae (Montane wood mouse)
  • H. endorobae (Mount Kenya wood mouse)
  • H. grandis (Mount Oku hylomyscus)
  • H. heinrichorum (Heinrich's wood mouse)
  • H. kerbispeterhansi (Kerbis Peterhans's wood mouse)
  • H. pamfi (Dahomey Gap wood mouse)
  • H. parvus (Little wood mouse)
  • H. stella (Stella wood mouse)
  • H. vulcanorum (Albertine Rift wood mouse)
  • H. walterverheyeni (Walter Verheyen's mouse)
Eastern, central, and western Africa Size range: 5 cm (2 in) long, plus 5 cm (2 in) tail (Dahomey Gap wood mouse) to 12 cm (5 in) long, plus 15 cm (6 in) tail (Baer's wood mouse)[322]

Habitats: Forest[393]

Diets: Fruit and other vegetation, as well as animals[321]
Hyomys
(white-eared giant rat)
Thomas, 1904

Two species
  • H. dammermani (Western white-eared giant rat)
  • H. goliath (Eastern white-eared giant rat)
New Guinea Size range: 29 cm (11 in) long, plus 24 cm (9 in) tail (western white-eared giant rat) to 39 cm (15 in) long, plus 38 cm (15 in) tail (eastern white-eared giant rat)[322]

Habitats: Forest[394]

Diets: Shoots as well as other vegetation[321]
Hyorhinomys

Brown and white rat

Esselstyn, Achmadi, Handika, & Rowe, 2015

One species
  • H. stuempkei (Hog-nosed shrew rat)
Indonesia Size: 11–13 cm (4–5 in) long, plus 8–10 cm (3–4 in) tail[322]

Habitats: Forest[395]

Diet: Plant material and invertebrates[321]
Kadarsanomys Musser, 1981

One species
  • K. sodyi (Sody's tree rat)
Indonesia
Map of range
Size: 16–21 cm (6–8 in) long, plus 25–30 cm (10–12 in) tail[322]

Habitats: Forest[396]

Diet: Plant material and invertebrates[321]
Komodomys

Dead brown rat

Musser & Boeadi, 1980

One species
Indonesia Size: 12–18 cm (5–7 in) long, plus 12–17 cm (5–7 in) tail[322]

Habitats: Shrubland and forest[397]

Diet: Plant material and invertebrates[321]
Lamottemys Petter, 1986

One species
Cameroon Size: 13–15 cm (5–6 in) long, plus 12–14 cm (5–6 in) tail[322]

Habitats: Forest[398]

Diet: Plant material and invertebrates[321]
Leggadina
(short-tailed mouse)

Dead brown mouse

Thomas, 1910

Two species
  • L. forresti (Forrest's mouse)
  • L. lakedownensis (Lakeland Downs mouse, pictured)
Northern and central Australia Size range: 5 cm (2 in) long, plus 4 cm (2 in) tail (Lakeland Downs mouse) to 10 cm (4 in) long, plus 7 cm (3 in) tail (Forrest's mouse)[322]

Habitats: Desert, savanna, grassland, and shrubland[399]

Diets: Seeds and vegetation[321]
Lemniscomys
(striped grass mouse)

Brown and white mouse

Trouessart, 1881

Eleven species
  • L. barbarus (Barbary striped grass mouse, pictured)
  • L. bellieri (Bellier's striped grass mouse)
  • L. griselda (Griselda's striped grass mouse)
  • L. hoogstraali (Hoogstraal's striped grass mouse)
  • L. linulus (Senegal one-striped grass mouse)
  • L. macculus (Buffoon striped grass mouse)
  • L. mittendorfi (Mittendorf's striped grass mouse)
  • L. rosalia (Single-striped grass mouse)
  • L. roseveari (Rosevear's striped grass mouse)
  • L. striatus (Typical striped grass mouse)
  • L. zebra (Heuglin's striped grass mouse)
Sub-Saharan Africa Size range: 8 cm (3 in) long, plus 7 cm (3 in) tail (buffoon striped grass mouse) to 17 cm (7 in) long, plus 16 cm (6 in) tail (single-striped grass mouse)[322]

Habitats: Savanna, forest, grassland, and shrubland[400]

Diets: Grass, seeds, grains, and insects[321]
Lenomys

Drawing of brown and white rat

Thomas, 1898

One species
  • L. meyeri (Trefoil-toothed giant rat)
Indonesia Size: 23–30 cm (9–12 in) long, plus 24–30 cm (9–12 in) tail[322]

Habitats: Forest[401]

Diet: Plant material and invertebrates[321]
Lenothrix

Dead brown rat

Miller, 1903

One species
Malaysia and Brunei Size: 14–21 cm (6–8 in) long, plus 19–29 cm (7–11 in) tail[322]

Habitats: Forest[402]

Diet: Plant material and invertebrates[321]
Leopoldamys
(long-tailed giant rat)

Brown and white rat

Ellerman, 1947

Seven species
  • L. ciliatus (Sundaic mountain leopoldamys)
  • L. diwangkarai (Diwangkara's long-tailed giant rat)
  • L. edwardsi (Edwards's long-tailed giant rat)
  • L. milleti (Millet's leopoldamys)
  • L. neilli (Neill's long-tailed giant rat)
  • L. sabanus (Long-tailed giant rat, pictured)
  • L. siporanus (Mentawai long-tailed giant rat)
Southeastern Asia Size range: 17 cm (7 in) long, plus 22 cm (9 in) tail (Mentawai long-tailed giant rat) to 29 cm (11 in) long, plus 31 cm (12 in) tail (Edwards's long-tailed giant rat)[322]

Habitats: Forest[403]

Diets: Insects, other invertebrates, and a wide variety of vegetation[321]
Leporillus
(stick-nest rat)

Brown rat

Thomas, 1906

Two species
Southern and western Australia Size range: 17–26 cm (7–10 in) long, plus 14–18 cm (6–7 in) tail (greater stick-nest rat)[322]

Habitats: Savanna, grassland, desert, caves, rocky areas, and shrubland[404]

Diets: Succulents and other vegetation[321]
Leptomys
(narrow water rat)
Thomas, 1897

Five species
  • L. arfakensis (Arfak water rat)
  • L. elegans (Long-footed water rat)
  • L. ernstmayri (Ernst Mayr's water rat)
  • L. paulus (Small water rat)
  • L. signatus (Fly River water rat)
New Guinea Size range: 11 cm (4 in) long, plus 13 cm (5 in) tail (small water rat) to 20 cm (8 in) long, plus 17 cm (7 in) tail (long-footed water rat)[322]

Habitats: Forest[405]

Diets: Insects and small animals[321]
Limnomys
(mountain rat)

Dead brown rat

Mearns, 1905

Two species
Philippines Size range: 11 cm (4 in) long, plus 14 cm (6 in) tail (Mindanao mountain rat) to 15 cm (6 in) long, plus 18 cm (7 in) tail (Mindanao mountain rat)[322]

Habitats: Forest[406]

Diets: Plant material and invertebrates[321]
Lorentzimys Jentink, 1911

One species
New Guinea Size: 5–9 cm (2–4 in) long, plus 11–13 cm (4–5 in) tail[322]

Habitats: Forest[407]

Diet: Insects, vegetation, and fungi[321]
Macruromys
(small-toothed rat)
Stein, 1933

Two species
  • M. elegans (Lesser small-toothed rat)
  • M. major (Eastern small-toothed rat)
Western New Guinea and New Guinea Size range: 15 cm (6 in) long, plus 21 cm (8 in) tail (lesser small-toothed rat) to 26 cm (10 in) long, plus 34 cm (13 in) tail (eastern small-toothed rat)[322]

Habitats: Forest[408]

Diets: Vegetation[321]
Madromys

Brown and white rat

Sody, 1941

One species
  • M. blanfordi (Blanford's rat)
Southern Asia Size: 15–19 cm (6–7 in) long, plus 18–21 cm (7–8 in) tail[322]

Habitats: Caves, forest, and shrubland[409]

Diet: Plant material and invertebrates[321]
Malacomys
(swamp rat)

Brown and white rat

A. Milne-Edwards, 1877

Three species
  • M. cansdalei (Cansdale's swamp rat)
  • M. edwardsi (Edward's swamp rat)
  • M. longipes (Big-eared swamp rat, pictured)
Central and western Africa Size range: 12 cm (5 in) long, plus 15 cm (6 in) tail (Edward's swamp rat) to 18 cm (7 in) long, plus 20 cm (8 in) tail (big-eared swamp rat)[322]

Habitats: Forest, inland wetlands, and shrubland[410]

Diets: Fruit, seeds, nuts, roots, insects, slugs, snails, and crabs[321]
Mallomys
(woolly rat)

Gray rat

Thomas, 1898

Four species
  • M. aroaensis (De Vis's woolly rat)
  • M. gunung (Alpine woolly rat, pictured)
  • M. istapantap (Subalpine woolly rat)
  • M. rothschildi (Rothschild's woolly rat)
New Guinea Size range: 34 cm (13 in) long, plus 33 cm (13 in) tail (De Vis's woolly rat) to 47 cm (19 in) long, plus 37 cm (15 in) tail (alpine woolly rat)[322]

Habitats: Grassland, forest, caves, rocky areas, and shrubland[411]

Diets: Shoots as well as other vegetation[321]
Mammelomys
(large mosaic-tailed rat)
Menzies, 1996

Two species
  • M. lanosus (Large-scaled mosaic-tailed rat)
  • M. rattoides (Large mosaic-tailed rat)
New Guinea Size range: 13 cm (5 in) long, plus 10 cm (4 in) tail (large-scaled mosaic-tailed rat) to 22 cm (9 in) long, plus 15 cm (6 in) tail (large mosaic-tailed rat)[322]

Habitats: Forest[412]

Diets: Plant material and invertebrates[321]
Margaretamys
(margareta rat)

Dead brown rat

Musser, 1981

Four species
  • M. beccarii (Beccari's margareta rat, pictured)
  • M. christinae (Christine's margareta rat)
  • M. elegans (Elegant margareta rat)
  • M. parvus (Little margareta rat)
Indonesia Size range: 9 cm (4 in) long, plus 15 cm (6 in) tail (little margareta rat) to 20 cm (8 in) long, plus 29 cm (11 in) tail (elegant margareta rat)[322]

Habitats: Forest[413]

Diets: Plant material and invertebrates[321]
Mastacomys

Gray mouse

Thomas, 1882

One species
Southeastern Australia Size: 14–20 cm (6–8 in) long, plus 10–14 cm (4–6 in) tail[322]

Habitats: Forest, shrubland, grassland, and inland wetlands[414]

Diet: Plant material and invertebrates[321]
Mastomys
(multimammate mouse)

Brown and white mouse

Thomas, 1915

Eight species
  • M. awashensis (Awash multimammate mouse)
  • M. coucha (Southern multimammate mouse, pictured)
  • M. erythroleucus (Guinea multimammate mouse)
  • M. huberti (Hubert's multimammate mouse)
  • M. kollmannspergeri (Verheyen's multimammate mouse)
  • M. natalensis (Natal multimammate mouse)
  • M. pernanus (Dwarf multimammate mouse)
  • M. shortridgei (Shortridge's multimammate mouse)
Morocco and Sub-Saharan Africa Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (dwarf multimammate mouse) to 18 cm (7 in) long, plus 16 cm (6 in) tail (Guinea multimammate mouse)[322]

Habitats: Savanna, grassland, inland wetlands, forest, and shrubland[415]

Diets: Grass, seeds, and insects[321]
Maxomys
(spiny rat)

Brown and white rat

Sody, 1936

Eighteen species
  • M. alticola (Mountain spiny rat)
  • M. baeodon (Small Bornean maxomys)
  • M. bartelsii (Bartels's spiny rat)
  • M. dollmani (Dollman's spiny rat)
  • M. hellwaldii (Hellwald's spiny rat)
  • M. hylomyoides (Sumatran spiny rat)
  • M. inas (Malayan mountain spiny rat)
  • M. inflatus (Fat-nosed spiny rat)
  • M. moi (Mo's spiny rat)
  • M. musschenbroekii (Musschenbroek's spiny rat)
  • M. ochraceiventer (Chestnut-bellied spiny rat)
  • M. pagensis (Pagai spiny rat)
  • M. panglima (Palawan spiny rat)
  • M. rajah (Rajah spiny rat)
  • M. surifer (Red spiny rat)
  • M. tajuddinii (Tajuddin's spiny rat)
  • M. wattsi (Watts's spiny rat)
  • M. whiteheadi (Whitehead's spiny rat, pictured)
Southeastern Asia Size range: 9 cm (4 in) long, plus 10 cm (4 in) tail (Tajuddin's spiny rat) to 23 cm (9 in) long, plus 21 cm (8 in) tail (rajah spiny rat)[322]

Habitats: Forest and shrubland[416]

Diets: Roots, fruit, and other vegetation, as well as invertebrates and small vertebrates[321]
Melasmothrix Miller & Hollister, 1921

One species
  • M. naso (Sulawesian shrew rat)
Indonesia Size: 11–13 cm (4–5 in) long, plus 8–10 cm (3–4 in) tail[322]

Habitats: Forest[417]

Diet: Worms and insect larvae[321]
Melomys
(mosaic-tailed rat)
Thomas, 1922

22 species
  • M. aerosus (Dusky mosaic-tailed rat)
  • M. arcium (Rossel Island melomys)
  • M. bannisteri (Bannister's rat)
  • M. bougainville (Bougainville mosaic-tailed rat)
  • M. burtoni (Grassland mosaic-tailed rat)
  • M. capensis (Cape York melomys)
  • M. caurinus (Short-tailed Talaud mosaic-tailed rat)
  • M. cervinipes (Fawn-footed mosaic-tailed rat)
  • M. cooperae (Yamdena mosaic-tailed rat)
  • M. dollmani (Dollman's melomys)
  • M. fraterculus (Manusela mosaic-tailed rat)
  • M. frigicola (Snow Mountains grassland mosaic-tailed rat)
  • M. fulgens (Seram long-tailed mosaic-tailed rat)
  • M. howi (Riama mosaic-tailed rat)
  • M. leucogaster (White-bellied mosaic-tailed rat)
  • M. lutillus (Papua grassland mosaic-tailed rat)
  • M. matambuai (Manus Island mosaic-tailed rat)
  • M. obiensis (Obi mosaic-tailed rat)
  • M. paveli (Pavel's Seram mosaic-tailed rat)
  • M. rubicola (Bramble Cay melomys)
  • M. rufescens (Black-tailed mosaic-tailed rat)
  • M. talaudium (Long-tailed Talaud mosaic-tailed rat)
Southeastern Asia and Australia Size range: 9–20 cm (4–8 in) long, plus 10–18 cm (4–7 in) tail (fawn-footed mosaic-tailed rat)[322]

Habitats: Savanna, grassland, forest, inland wetlands, and shrubland[418]

Diets: Fruit, berries, and other vegetation[321]
Mesembriomys
(tree-rat)

Drawing of brown and white rat

Palmer, 1906

Two species
Northern Australia Size range: 18 cm (7 in) long, plus 29 cm (11 in) tail (golden-backed tree-rat) to 30 cm (12 in) long, plus 41 cm (16 in) tail (black-footed tree-rat)[322]

Habitats: Savanna and forest[419]

Diets: Seeds, nuts, and insects[321]
Micaelamys
(rock mouse)

Brown rat

Ellerman, 1941

Two species
  • M. granti (Grant's rock mouse)
  • M. namaquensis (Namaqua rock rat, pictured)
Southern Africa Size range: 8–15 cm (3–6 in) long, plus 10–20 cm (4–8 in) tail (Namaqua rock rat)[322]

Habitats: Savanna, grassland, desert, forest, rocky areas, and shrubland[420]

Diets: Grain, seeds, roots, nuts, and fruit[321]
Microhydromys
(groove-toothed shrew mouse)
Tate & Archbold, 1941

Two species
  • M. argenteus (Southern groove-toothed moss mouse)
  • M. richardsoni (Northern groove-toothed shrew mouse)
New Guinea Size range: 7–9 cm (3–4 in) long, plus 8–10 cm (3–4 in) tail (northern groove-toothed shrew mouse)[322]

Habitats: Savanna and forest[421]

Diets: Insects[321]
Micromys

Brown mouse

Dehne, 1841

One species
Europe and Asia
Map of range
Size: 4–8 cm (2–3 in) long, plus 3–8 cm (1–3 in) tail[322]

Habitats: Forest and inland wetlands[422]

Diet: Seeds, vegetation, and insects, as well as bird eggs[321]
Millardia
(soft-furred rat)

Brown rat

Thomas, 1911

Four species
  • M. gleadowi (Sand-colored soft-furred rat, pictured)
  • M. kathleenae (Miss Ryley's soft-furred rat)
  • M. kondana (Kondana rat)
  • M. meltada (Soft-furred rat)
Southern Asia Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (sand-colored soft-furred rat) to 20 cm (8 in) long, plus 19 cm (7 in) tail (Kondana rat)[322]

Habitats: Grassland, desert, forest, rocky areas, and shrubland[423]

Diets: Grain, seeds, and swamp vegetation[321]
Mirzamys
(moss rat)
Helgen & Helgen, 2009

Two species
  • M. louiseae (Mirza's western moss rat)
  • M. norahae (Mirza's eastern moss rat)
New Guinea Size range: 9 cm (4 in) long, plus 10 cm (4 in) tail (Mirza's western moss rat) to 12 cm (5 in) long, plus 10 cm (4 in) tail (Mirza's eastern moss rat)[322]

Habitats: Forest and grassland[424]

Diets: Plant material and invertebrates[321]
Muriculus Rüppell, 1842

One species
  • M. imberbis (Ethiopian striped mouse)
Ethiopia Size: 7–8 cm (3 in) long, plus 4–6 cm (2 in) tail[322]

Habitats: Grassland[425]

Diet: Plant material and invertebrates[321]
Mus
(mouse)

Brown and white mouse

Linnaeus, 1758

39 species
  • M. baoulei (Baoule's mouse)
  • M. booduga (Little Indian field mouse)
  • M. bufo (Toad mouse)
  • M. callewaerti (Callewaert's mouse)
  • M. caroli (Ryukyu mouse)
  • M. cervicolor (Fawn-colored mouse)
  • M. cookii (Cook's mouse)
  • M. crociduroides (Sumatran shrewlike mouse)
  • M. cypriacus (Cypriot mouse)
  • M. famulus (Servant mouse)
  • M. fernandoni (Ceylon spiny mouse)
  • M. fragilicauda (Sheath-tailed mouse)
  • M. goundae (Gounda mouse)
  • M. harennensis (Harenna mouse)
  • M. haussa (Hausa mouse)
  • M. indutus (Desert pygmy mouse)
  • M. macedonicus (Macedonian mouse)
  • M. mahomet (Mahomet mouse)
  • M. mattheyi (Matthey's mouse)
  • M. mayori (Mayor's mouse)
  • M. minutoides (African pygmy mouse, pictured)
  • M. musculoides (Temminck's mouse)
  • M. musculus (House mouse)
  • M. neavei (Neave's mouse)
  • M. oubanguii (Oubangui mouse)
  • M. pahari (Gairdner's shrewmouse)
  • M. phillipsi (Phillips's mouse)
  • M. platythrix (Stone-loving mouse)
  • M. saxicola (Rock-loving mouse)
  • M. setulosus (Peters's mouse)
  • M. setzeri (Setzer's pygmy mouse)
  • M. shortridgei (Shortridge's mouse)
  • M. sorella (Thomas's pygmy mouse)
  • M. spicilegus (Steppe mouse)
  • M. spretus (Algerian mouse)
  • M. tenellus (Delicate mouse)
  • M. terricolor (Earth-colored mouse)
  • M. triton (Gray-bellied pygmy mouse)
  • M. vulcani (Volcano mouse)
Africa, Europe, and Asia Size range: 4 cm (2 in) long, plus 2 cm (0.8 in) tail (African pygmy mouse) to 13 cm (5 in) long, plus 9 cm (4 in) tail (stone-loving mouse)[322]

Habitats: Grassland[426]

Diets: Seeds, roots, leaves, stems, and other vegetation, as well as insects and some meat[321]
Musseromys
(tree-mouse)
Heaney, Balete, Rickart, Veluz, & Jansa, 2009

Four species
  • M. anacuao (Sierra Madre tree-mouse)
  • M. beneficus (Mount Pulag tree-mouse)
  • M. gulantang (Banahaw tree mouse)
  • M. inopinatus (Amuyao tree-mouse)
Philippines Size range: 7–9 cm (3–4 in) long, plus 8 cm (3 in) tail (Mount Pulag tree-mouse)[322]

Habitats: Forest[427]

Diets: Plant material and invertebrates[321]
Mylomys
(groove-toothed rat)
Thomas, 1906

Two species
  • M. dybowskii (African groove-toothed rat)
  • M. rex (Ethiopian mylomys)
Northern Sub-Saharan Africa Size range: 12 cm (5 in) long, plus 10 cm (4 in) tail (African groove-toothed rat) to 21 cm (8 in) long, plus 18 cm (7 in) tail (Ethiopian mylomys)[322]

Habitats: Savanna and forest[428]

Diets: Grass and leaves[321]
Myomyscus
(white-footed rat)

Brown mouse

Shortridge, 1942

Four species
  • M. angolensis (Myomyscus angolensis)
  • M. brockmani (Brockman's rock mouse)
  • M. verreauxii (Verreaux's mouse, pictured)
  • M. yemeni (Yemeni mouse)
Sub-Saharan Africa, southern Saudi Arabia and Yemen Size range: 7 cm (3 in) long, plus 15 cm (6 in) tail (Yemeni mouse) to 13 cm (5 in) long, plus 18 cm (7 in) tail (Brockman's rock mouse)[322]

Habitats: Savanna, rocky areas, and shrubland[429]

Diets: Insects, leaves, and shoots[321]
Nesokia
(short-tailed bandicoot rat)

Drawing of brown rat

Gray, 1842

Two species
Scattered Asia and Egypt Size range: 15 cm (6 in) long, plus 9 cm (4 in) tail (short-tailed bandicoot rat) to 26 cm (10 in) long, plus 27 cm (11 in) tail (Bunn's short-tailed bandicoot rat)[322]

Habitats: Forest, inland wetlands, shrubland, and grassland[430]

Diets: Grass, grain, roots, fruit, and vegetables[321]
Nesoromys Thomas, 1922

One species
  • N. ceramicus (Ceram rat)
Seram Island in Indonesia Size: 11–14 cm (4–6 in) long, plus 12–14 cm (5–6 in) tail[322]

Habitats: Forest[431]

Diet: Plant material and invertebrates[321]
Nilopegamys

Drawing of brown and white rat

Osgood, 1928

One species
Ethiopia
Map of range
Size: About 15 cm (6 in)[322]

Habitats: Inland wetlands[432]

Diet: Plant material and invertebrates[321]
Niviventer
(white-bellied rat)

Brown and white rat

Marshall, 1976

Seventeen species
  • N. andersoni (Anderson's white-bellied rat)
  • N. brahma (Brahma white-bellied rat)
  • N. cameroni (Cameron Highlands white-bellied rat)
  • N. confucianus (Chinese white-bellied rat)
  • N. coninga (Coxing's white-bellied rat)
  • N. cremoriventer (Dark-tailed tree rat)
  • N. culturatus (Oldfield white-bellied rat)
  • N. eha (Smoke-bellied rat)
  • N. excelsior (Large white-bellied rat)
  • N. fraternus (Montane Sumatran white-bellied rat)
  • N. fulvescens (Chestnut white-bellied rat)
  • N. hinpoon (Limestone rat)
  • N. langbianis (Lang Bian white-bellied rat)
  • N. lepturus (Narrow-tailed white-bellied rat)
  • N. niviventer (White-bellied rat, pictured)
  • N. rapit (Long-tailed mountain rat)
  • N. tenaster (Tenasserim white-bellied rat)
Eastern, southern, and southeastern Asia Size range: 10 cm (4 in) long, plus 15 cm (6 in) tail (chestnut white-bellied rat) to 21 cm (8 in) long, plus 19 cm (7 in) tail (Chinese white-bellied rat)[322]

Habitats: Forest and shrubland[433]

Diets: Plant material and invertebrates[321]
Notomys
(hopping mouse)

Brown mouse

Lesson, 1842

Ten species
Australia Size range: 7 cm (3 in) long, plus 11 cm (4 in) tail (dusky hopping mouse) to 13 cm (5 in) long, plus 16 cm (6 in) tail (Mitchell's hopping mouse)[322]

Habitats: Savanna, grassland, desert, forest, rocky areas, and shrubland[434]

Diets: Berries, leaves, seeds, and other vegetation[321]
Oenomys
(rufous-nosed rat)

Brown rat

Thomas, 1904

Two species
  • O. hypoxanthus (Common rufous-nosed rat, pictured)
  • O. ornatus (Ghana rufous-nosed rat)
Central and western Africa Size range: 13–19 cm (5–7 in) long, plus 15–20 cm (6–8 in) tail (common rufous-nosed rat)[322]

Habitats: Savanna, forest, grassland, and shrubland[435]

Diets: Vegetation, as well as insects[321]
Otomys
(vlei rat)

Brown rat

F. Cuvier, 1824

28 species
  • O. anchietae (Angolan vlei rat)
  • O. angoniensis (Angoni vlei rat)
  • O. auratus (South East African vlei rat)
  • O. barbouri (Barbour's vlei rat)
  • O. burtoni (Burton's vlei rat)
  • O. cheesmani (Cheesman's vlei rat)
  • O. cuanzensis (Cuanza vlei rat)
  • O. dartmouthi (Ruwenzori vlei rat)
  • O. denti (Dent's vlei rat)
  • O. dollmani (Dollman's vlei rat)
  • O. fortior (Charada vlei rat)
  • O. helleri (Heller's vlei rat)
  • O. irroratus (Southern African vlei rat)
  • O. jacksoni (Mount Elgon vlei rat)
  • O. karoensis (Fynbos vlei rat)
  • O. lacustris (Tanzanian vlei rat)
  • O. laminatus (Laminate vlei rat)
  • O. occidentalis (Western vlei rat)
  • O. orestes (Afroalpine vlei rat, pictured)
  • O. simiensis (Simien vlei rat)
  • O. sloggetti (Sloggett's vlei rat)
  • O. thomasi (Thomas's vlei rat)
  • O. tropicalis (Tropical vlei rat)
  • O. typus (Ethiopian vlei rat)
  • O. unisulcatus (Karoo vlei rat)
  • O. uzungwensis (Uzungwe vlei rat)
  • O. yaldeni (Yalden's vlei rat)
  • O. zinki (Mount Kilimanjaro vlei rat)
Sub-Saharan Africa Size range: 11 cm (4 in) long, plus 8 cm (3 in) tail (Fynbos vlei rat) to 22 cm (9 in) long, plus 13 cm (5 in) tail (Angolan vlei rat)[322]

Habitats: Savanna, grassland, forest, inland wetlands, rocky areas, and shrubland[436]

Diets: Grass, semiaquatic plants, and shoots, as well as grain, seeds, berries, roots, and bark[321]
Palawanomys Musser & Newcomb, 1983

One species
  • P. furvus (Palawan soft-furred mountain rat)
Philippines
Map of range
Size: 13–16 cm (5–6 in) long, plus 14–17 cm (6–7 in) tail[322]

Habitats: Forest[437]

Diet: Plant material and invertebrates[321]
Papagomys

Dead brown rat

Sody, 1941

One species
Indonesia Size: 27–43 cm (11–17 in) long, plus 26–36 cm (10–14 in) tail[322]

Habitats: Forest[438]

Diet: Unknown[321]
Parahydromys Poche, 1906

One species
  • P. asper (New Guinea waterside rat)
New Guinea
Map of range
Size: 21–23 cm (8–9 in) long, plus 23–27 cm (9–11 in) tail[322]

Habitats: Inland wetlands and forest[439]

Diet: Insects and other invertebrates[321]
Paraleptomys
(short-haired water rat)
Tate & Archbold, 1941

Two species
  • P. rufilatus (Northern water rat)
  • P. wilhelmina (Short-haired water rat)
New Guinea
Map of range
Size range: 10 cm (4 in) long, plus 12 cm (5 in) tail (short-haired water rat) to 14 cm (6 in) long, plus 15 cm (6 in) tail (northern water rat)[322]

Habitats: Forest[440]

Diets: Plant material and invertebrates[321]
Paramelomys
(long-nosed mosaic-tailed rat)

Dead brown rat

Rümmler, 1936

Nine species
  • P. gressitti (Gressitt's mosaic-tailed rat)
  • P. levipes (Long-nosed mosaic-tailed rat)
  • P. lorentzii (Lorentz's mosaic-tailed rat)
  • P. mollis (Thomas's mosaic-tailed rat)
  • P. moncktoni (Moncton's mosaic-tailed rat)
  • P. naso (Long-nosed paramelomys)
  • P. platyops (Lowland mosaic-tailed rat, pictured)
  • P. rubex (Mountain mosaic-tailed rat)
  • P. steini (Stein's paramelomys)
New Guinea and nearby islands Size range: 10 cm (4 in) long, plus 9 cm (4 in) tail (mountain mosaic-tailed rat) to 20 cm (8 in) long, plus 13 cm (5 in) tail (long-nosed paramelomys)[322]

Habitats: Savanna and forest[441]

Diets: Fruit, berries, and other vegetation[321]
Parotomys
(whistling rat)

Brown rat

Thomas, 1918

Two species
  • P. brantsii (Brants's whistling rat, pictured)
  • P. littledalei (Littledale's whistling rat)
Southern Africa Size range: 12–17 cm (5–7 in) long, plus 8–11 cm (3–4 in) tail (Littledale's whistling rat)[322]

Habitats: Desert and shrubland[442]

Diets: Grass, seeds, and shoots[321]
Paucidentomys Rowe, Achmadi, & Esselstyn, 2012

One species
Indonesia
Map of range
Size: 15–17 cm (6–7 in) long, plus 19–20 cm (7–8 in) tail[322]

Habitats: Forest[443]

Diet: Plant material and invertebrates[321]
Paulamys Musser, 1986

One species
  • P. naso (Paulamys)
Indonesia Size: About 16 cm (6 in)[322]

Habitats: Forest[444]

Diet: Insects, snails, worms, fungi, and fruit[321]
Pelomys
(groove-toothed swamp rat)
Peters, 1852

Five species
  • P. campanae (Bell groove-toothed swamp rat)
  • P. fallax (Creek groove-toothed swamp rat)
  • P. hopkinsi (Hopkins's groove-toothed swamp rat)
  • P. isseli (Issel's groove-toothed swamp rat)
  • P. minor (Least groove-toothed swamp rat)
Central Africa Size range: 10 cm (4 in) long, plus 10 cm (4 in) tail (least groove-toothed swamp rat) to 17 cm (7 in) long, plus 15 cm (6 in) tail (bell groove-toothed swamp rat)[322]

Habitats: Savanna, inland wetlands, and grassland[445]

Diets: Grass, swamp vegetation, and grains[321]
Phloeomys
(slender-tailed cloud rat)

Gray and black rat

Waterhouse, 1839

Two species
Philippines Size range: 39 cm (15 in) long, plus 32 cm (13 in) tail (northern Luzon giant cloud rat) to 44 cm (17 in) long, plus 31 cm (12 in) tail (southern giant slender-tailed cloud rat)[322]

Habitats: Forest and shrubland[446]

Diets: Vegetation[321]
Pithecheir
(tree rat)
Lesson, 1840

Two species
  • P. melanurus (Red tree rat)
  • P. parvus (Malayan tree rat)
Malaysia and Indonesia Size range: 12–18 cm (5–7 in) long, plus 15–22 cm (6–9 in) tail (Malayan tree rat)[322]

Habitats: Forest[447]

Diets: Vegetation and insects[321]
Pithecheirops Emmons, 1993

One species
Eastern Malaysia Size: About 11 cm (4 in)[322]

Habitats: Forest[448]

Diet: Plant material and invertebrates[321]
Pogonomelomys
(lowland brush mouse)

Drawing of brown and white mouse

Rümmler, 1936

Three species
  • P. brassi (Grey pogonomelomys)
  • P. bruijni (Lowland brush mouse, pictured)
  • P. mayeri (Shaw Mayer's brush mouse)
New Guinea Size range: 13 cm (5 in) long, plus 15 cm (6 in) tail (Shaw Mayer's brush mouse) to 18 cm (7 in) long, plus 20 cm (8 in) tail (grey pogonomelomys)[322]

Habitats: Forest[449]

Diets: Plant material and invertebrates[321]
Pogonomys
(prehensile-tailed rat)

Gray mouse

A. Milne-Edwards, 1877

Five species
  • P. championi (Champion's tree mouse)
  • P. fergussoniensis (D'Entrecasteaux Archipelago pogonomys)
  • P. loriae (Large tree mouse)
  • P. macrourus (Chestnut tree mouse, pictured)
  • P. sylvestris (Gray-bellied tree mouse)
Northern Australia and New Guinea and nearby islands Size range: 9 cm (4 in) long, plus 12 cm (5 in) tail (chestnut tree mouse) to 20 cm (8 in) long, plus 26 cm (10 in) tail (D'Entrecasteaux Archipelago pogonomys)[322]

Habitats: Forest[450]

Diets: Leaves and grass and bamboo shoots[321]
Praomys
(soft-furred mouse)

Gray mouse

Thomas, 1915

Seventeen species
  • P. coetzeei (Coetzee's soft-furred mouse)
  • P. daltoni (Dalton's mouse, pictured)
  • P. degraaffi (De Graaff's soft-furred mouse)
  • P. delectorum (Delectable soft-furred mouse)
  • P. derooi (Deroo's mouse)
  • P. hartwigi (Hartwig's soft-furred mouse)
  • P. jacksoni (Jackson's soft-furred mouse)
  • P. lukolelae (Lukolela swamp rat)
  • P. minor (Least soft-furred mouse)
  • P. misonnei (Misonne's soft-furred mouse)
  • P. morio (Cameroon soft-furred mouse)
  • P. mutoni (Muton's soft-furred mouse)
  • P. obscurus (Gotel Mountain soft-furred mouse)
  • P. petteri (Petter's soft-furred mouse)
  • P. rostratus (Forest soft-furred mouse)
  • P. tullbergi (Tullberg's soft-furred mouse)
  • P. verschureni (Verschuren's swamp rat)
Sub-Saharan Africa Size range: 8 cm (3 in) long, plus 10 cm (4 in) tail (least soft-furred mouse) to 15 cm (6 in) long, plus 18 cm (7 in) tail (forest soft-furred mouse)[322]

Habitats: Savanna, grassland, inland wetlands, forest, and rocky areas[451]

Diets: Fruit, seeds, vegetation, and insects[321]
Protochromys Menzies, 1996

One species
  • P. fellowsi (Red-bellied mosaic-tailed rat)
Papua New Guinea Size: 13–18 cm (5–7 in) long, plus 18–21 cm (7–8 in) tail[322]

Habitats: Forest[452]

Diet: Fruit, berries, and other vegetation[321]
Pseudohydromys
(moss mouse)
Rümmler, 1934

Twelve species
  • P. berniceae (Bishop's moss mouse)
  • P. carlae (Huon small-toothed moss mouse)
  • P. elanorae (Laurie's moss mouse)
  • P. ellermani (One-toothed shrew mouse)
  • P. fuscus (Mottled-tailed shrew mouse)
  • P. germani (German's one-toothed moss mouse)
  • P. murinus (Eastern shrew mouse)
  • P. musseri (Musser's shrew mouse)
  • P. occidentalis (Western shrew mouse)
  • P. patriciae (Woolley's moss mouse)
  • P. pumehanae (Southern small-toothed moss mouse)
  • P. sandrae (White-bellied moss mouse)
New Guinea Size range: 7 cm (3 in) long, plus 7 cm (3 in) tail (Laurie's moss mouse) to 12 cm (5 in) long, plus 10 cm (4 in) tail (western shrew mouse)[322]

Habitats: Forest, grassland, and shrubland[453]

Diets: Insects[321]
Pseudomys
(pebble-mound mouse)

Brown mouse

Gray, 1832

23 species
Australia and Papua New Guinea Size range: 5 cm (2 in) long, plus 7 cm (3 in) tail (Bolam's mouse) to 16 cm (6 in) long, plus 14 cm (6 in) tail (Hastings River mouse)[322]

Habitats: Savanna, grassland, desert, forest, inland wetlands, rocky areas, and shrubland[454]

Diets: Seeds, roots, other vegetation, and insects[321]
Rattus
(rat)

Gray rat

Fischer von Waldheim, 1803

68 species
  • R. adustus (Sunburned rat)
  • R. andamanensis (Sikkim rat)
  • R. annandalei (Annandale's rat)
  • R. arfakiensis (Vogelkop mountain rat)
  • R. argentiventer (Ricefield rat)
  • R. arrogans (Western New Guinea mountain rat)
  • R. baluensis (Summit rat)
  • R. blangorum (Aceh rat)
  • R. bontanus (Bonthain rat)
  • R. burrus (Nonsense rat)
  • R. colletti (Dusky rat)
  • R. detentus (Manus Island spiny rat)
  • R. elaphinus (Sula rat)
  • R. enganus (Enggano rat)
  • R. everetti (Philippine forest rat)
  • R. exulans (Polynesian rat)
  • R. facetus (Lore Lindu xanthurus rat)
  • R. feliceus (Spiny Ceram rat)
  • R. fuscipes (Bush rat)
  • R. giluwensis (Giluwe rat)
  • R. hainaldi (Hainald's Flores Island Rat)
  • R. hoffmanni (Hoffmann's rat)
  • R. hoogerwerfi (Hoogerwerf's rat)
  • R. jobiensis (Japen rat)
  • R. koopmani (Koopman's rat)
  • R. korinchi (Korinch's rat)
  • R. leucopus (Cape York rat)
  • R. losea (Lesser ricefield rat)
  • R. lugens (Mentawai rat)
  • R. lutreolus (Australian swamp rat)
  • R. macleari (Maclear's rat)
  • R. marmosurus (Opossum rat)
  • R. mindorensis (Mindoro black rat)
  • R. mollicomulus (Little soft-furred rat)
  • R. montanus (Nillu rat)
  • R. mordax (Eastern rat)
  • R. morotaiensis (Moluccan prehensile-tailed rat)
  • R. nativitatis (Bulldog rat)
  • R. niobe (Moss-forest rat)
  • R. nitidus (Himalayan field rat)
  • R. norvegicus (Brown rat, pictured)
  • R. novaeguineae (New Guinean rat)
  • R. omichlodes (Arianus's rat)
  • R. osgoodi (Osgood's rat)
  • R. palmarum (Palm rat)
  • R. pelurus (Peleng rat)
  • R. pococki (Pocock's highland rat)
  • R. praetor (Large New Guinea spiny rat)
  • R. pyctoris (Turkestan rat)
  • R. ranjiniae (Kerala rat)
  • R. rattus (Black rat)
  • R. richardsoni (Glacier rat)
  • R. salocco (Southeastern xanthurus rat)
  • R. sanila (New Ireland forest rat)
  • R. satarae (Sahyadris forest rat)
  • R. simalurensis (Simalur rat)
  • R. sordidus (Dusky field rat)
  • R. steini (Stein's rat)
  • R. stoicus (Andaman rat)
  • R. tanezumi (Tanezumi rat)
  • R. tawitawiensis (Tawitawi forest rat)
  • R. timorensis (Timor rat)
  • R. tiomanicus (Malayan field rat)
  • R. tunneyi (Pale field rat)
  • R. vandeuseni (Van Deusen's rat)
  • R. verecundus (Slender rat)
  • R. villosissimus (Long-haired rat)
  • R. xanthurus (Yellow-tailed rat)
Europe, Asia, Africa, and Australia Size range: 6 cm (2 in) long, plus 7 cm (3 in) tail (dusky rat) to 29 cm (11 in) long, plus 25 cm (10 in) tail (brown rat)[322]

Habitats: Coastal marine, savanna, grassland, desert, intertidal marine, inland wetlands, forest, rocky areas, and shrubland[455]

Diets: A wide variety of plant, insects, and animals[321]
Rhabdomys
(four-striped grass rat)

Brown rat

Thomas, 1916

Four species
  • R. bechuanae (West-central four-striped grass rat, pictured)
  • R. dilectus (Mesic four-striped grass rat)
  • R. intermedius (Karoo four-striped grass rat)
  • R. pumilio (Four-striped grass mouse)
Sub-Saharan Africa Size range: 9–14 cm (4–6 in) long, plus 9–13 cm (4–5 in) tail (four-striped grass mouse)[322]

Habitats: Savanna, grassland, and shrubland[456]

Diets: Roots, seeds, berries, and grains, as well as insects and eggs[321]
Rhynchomys
(shrewlike rat)

Drawing of brown rats

Thomas, 1895

Four species
  • R. banahao (Banahao shrew-rat)
  • R. isarogensis (Isarog shrew-rat)
  • R. soricoides (Mount Data shrew-rat, pictured)
  • R. tapulao (Tapulao shrew-rat)
Philippines Size range: 16 cm (6 in) long, plus 12 cm (5 in) tail (Tapulao shrew-rat) to 20 cm (8 in) long, plus 17 cm (7 in) tail (Mount Data shrew-rat)[322]

Habitats: Forest[457]

Diets: Insects and worms[321]
Saxatilomys

Gray rat

Musser, 2005

One species
Indonesia
Map of range
Size: 15–17 cm (6–7 in) long, plus 18–20 cm (7–8 in) tail[322]

Habitats: Forest and rocky areas[458]

Diet: Plant material and invertebrates[321]
Solomys
(naked-tailed rat)
Thomas, 1922

Four species
  • S. ponceleti (Poncelet's giant rat)
  • S. salamonis (Ugi naked-tailed rat)
  • S. salebrosus (Bougainville naked-tailed rat)
  • S. sapientis (Isabel naked-tailed rat)
Solomon Islands and Papua New Guinea Size range: 18 cm (7 in) long, plus 19 cm (7 in) tail (Isabel naked-tailed rat) to 33 cm (13 in) long, plus 25 cm (10 in) tail (Bougainville naked-tailed rat)[322]

Habitats: Forest[459]

Diets: Coconuts and nuts[321]
Sommeromys Musser & Durden, 2002

One species
  • S. macrorhinos (Sommer's Sulawesi rat)
Indonesia Size: 9–11 cm (4 in) long, plus 17–20 cm (7–8 in) tail[322]

Habitats: Forest[460]

Diet: Plant material and invertebrates[321]
Soricomys
(shrew mouse)

Brown mouse on stamp

Balete, Rickart, Heaney, Alviola, Duya, Duya, Sosa, & Jansa, 2012

Four species
  • S. kalinga (Kalinga shrew mouse, pictured)
  • S. leonardicoi (Co's shrew mouse)
  • S. montanus (Southern Cordillera shrew mouse)
  • S. musseri (Sierra Madre shrew mouse)
Philippines Size range: 9–12 cm (4–5 in) long, plus 8–10 cm (3–4 in) tail (Co's shrew mouse)[322]

Habitats: Forest[461]

Diets: Worms and soft-bodied invertebrates[321]
Srilankamys Musser, 1981

One species
  • S. ohiensis (Ohiya rat)
Sri Lanka
Map of range
Size: About 15 cm (6 in)[322]

Habitats: Forest[462]

Diet: Plant material and invertebrates[321]
Stenocephalemys
(narrow-headed rat)

Brown rat

Frick, 1914

Four species
  • S. albipes (Ethiopian white-footed mouse)
  • S. albocaudata (Ethiopian narrow-headed rat, pictured)
  • S. griseicauda (Gray-tailed narrow-headed rat)
  • S. ruppi (Rupp's mouse)
Ethiopia Size range: 10–20 cm (4–8 in) long, plus 11–18 cm (4–7 in) tail (Ethiopian narrow-headed rat)[322]

Habitats: Forest, grassland, and shrubland[463]

Diets: Insects, leaves, and shoots[321]
Stochomys Thomas, 1926

One species
Central Africa Size: 10–17 cm (4–7 in) long, plus 18–23 cm (7–9 in) tail[322]

Habitats: Forest[464]

Diet: Fruit, as well as vegetation and insects[321]
Sundamys
(giant Sunda rat)
Musser & Newcomb, 1983

Three species
  • S. infraluteus (Mountain giant Sunda rat)
  • S. maxi (Bartels's rat)
  • S. muelleri (Müller's giant Sunda rat)
Indonesia and Malaysia, Southeastern Asia, and Indonesia Size range: 18–30 cm (7–12 in) long, plus 21–37 cm (8–15 in) tail (Müller's giant Sunda rat)[322]

Habitats: Forest and shrubland[465]

Diets: Fruit, leaves, shoots, insects, crabs, snails, and lizards[321]
Taeromys
(Sulawesi rat)

Drawing of brown and white rats

Sody, 1841

Eight species
  • T. arcuatus (Salokko rat)
  • T. callitrichus (Lovely-haired rat, pictured)
  • T. celebensis (Celebes rat)
  • T. dominator (Giant Sulawesi rat)
  • T. hamatus (Sulawesi montane rat)
  • T. microbullatus (Small-eared rat)
  • T. punicans (Sulawesi forest rat)
  • T. taerae (Tondano rat)
Indonesia Size range: 18 cm (7 in) long, plus 15 cm (6 in) tail (Sulawesi forest rat) to 28 cm (11 in) long, plus 33 cm (13 in) tail (giant Sulawesi rat)[322]

Habitats: Forest[466]

Diets: Fruit; Fruit, leaves, and insects[321]
Tarsomys
(long-footed rat)
Mearns, 1905

Two species
  • T. apoensis (Long-footed rat)
  • T. echinatus (Spiny long-footed rat)
Philippines Size range: 13 cm (5 in) long, plus 11 cm (4 in) tail (long-footed rat) to 18 cm (7 in) long, plus 16 cm (6 in) tail (spiny long-footed rat)[322]

Habitats: Forest[467]

Diets: Invertebrates[321]
Tateomys
(long-tailed shrew rat)
Musser, 1969

Two species
  • T. macrocercus (Long-tailed shrew rat)
  • T. rhinogradoides (Tate's shrew rat)
Indonesia Size range: 11 cm (4 in) long, plus 16 cm (6 in) tail (long-tailed shrew rat) to 16 cm (6 in) long, plus 17 cm (7 in) tail (Tate's shrew rat)[322]

Habitats: Forest[468]

Diets: Worms[321]
Thallomys
(acacia rat)

Gray rats

Thomas, 1920

Four species
  • T. loringi (Loring's rat)
  • T. nigricauda (Black-tailed tree rat, pictured)
  • T. paedulcus (Acacia rat)
  • T. shortridgei (Shortridge's rat)
Sub-Saharan Africa Size range: 11–17 cm (4–7 in) long, plus 12–20 cm (5–8 in) tail (black-tailed tree rat)[322]

Habitats: Savanna, forest, and shrubland[469]

Diets: Buds, leaves, and seeds, as well as berries, roots, and insects[321]
Thamnomys
(thicket rat)
Thomas, 1907

Four species
  • T. kempi (Kemp's thicket rat)
  • T. major (Hatt's thicket rat)
  • T. schoutedeni (Schouteden's thicket rat)
  • T. venustus (Charming thicket rat)
Central Africa Size range: 11 cm (4 in) long, plus 17 cm (7 in) tail (Schouteden's thicket rat) to 18 cm (7 in) long, plus 23 cm (9 in) tail (Kemp's thicket rat)[322]

Habitats: Forest[470]

Diets: Leaves and seeds[321]
Tokudaia
(Ryūkyū spiny rat)

Stuffed brown rat

Kuroda, 1943

Three species
  • T. muenninki (Muennink's spiny rat)
  • T. osimensis (Ryukyu spiny rat, pictured)
  • T. tokunoshimensis (Tokunoshima spiny rat)
Southern islands in Japan Size range: 10 cm (4 in) long, plus 8 cm (3 in) tail (Ryukyu spiny rat) to 18 cm (7 in) long, plus 14 cm (6 in) tail (Muennink's spiny rat)[322]

Habitats: Forest[471]

Diets: Plant material and invertebrates[321]
Tonkinomys Musser, Lunde, & Son, 2006

One species
Vietnam Size: 18–22 cm (7–9 in) long, plus 15–19 cm (6–7 in) tail[322]

Habitats: Forest, inland wetlands, and rocky areas[472]

Diet: Plant material and invertebrates[321]
Tryphomys Miller, 1910

One species
  • T. adustus (Luzon short-nosed rat)
Philippines
Map of range
Size: 17–20 cm (7–8 in) long, plus 15–19 cm (6–7 in) tail[322]

Habitats: Forest and grassland[473]

Diet: Plant material and invertebrates[321]
Uromys
(giant rat)

Brown and white rat

Peters, 1867

Eleven species
  • U. anak (Giant naked-tailed rat)
  • U. boeadii (Biak giant rat)
  • U. caudimaculatus (Giant white-tailed rat, pictured)
  • U. emmae (Emma's giant rat)
  • U. hadrourus (Masked white-tailed rat)
  • U. imperator (Emperor rat)
  • U. neobritannicus (Bismarck giant rat)
  • U. porculus (Guadalcanal rat)
  • U. rex (King rat)
  • U. siebersi (Great Key Island giant rat)
  • U. vika (Vangunu giant rat)
Southeastern Asia and northern Australia Size range: 17 cm (7 in) long, plus 18 cm (7 in) tail (masked white-tailed rat) to 38 cm (15 in) long, plus 36 cm (14 in) tail (giant white-tailed rat)[322]

Habitats: Inland wetlands, forest, and caves[474]

Diets: Coconuts, nuts, fruit, and flowers[321]
Vandeleuria
(long-tailed climbing mouse)

Brown and white mouse

Gray, 1842

Three species
  • V. nilagirica (Nilgiri long-tailed tree mouse)
  • V. nolthenii (Nolthenius's long-tailed climbing mouse)
  • V. oleracea (Asiatic long-tailed climbing mouse, pictured)
Southern and southeastern Asia Size range: 7 cm (3 in) long, plus 11 cm (4 in) tail (Asiatic long-tailed climbing mouse) to 13 cm (5 in) long, plus 13 cm (5 in) tail (Nolthenius's long-tailed climbing mouse)[322]

Habitats: Forest, grassland, and shrubland[475]

Diets: Fruit, buds, and shoorts[321]
Vernaya Anthony, 1941

One species
China
Map of range
Size: 5–8 cm (2–3 in) long, plus 10–14 cm (4–6 in) tail[322]

Habitats: Forest[476]

Diet: Plant material and invertebrates[321]
Waiomys

Brown rat

Rowe, Achmadi, & Esselstyn, 2014

One species
  • W. mamasae (Sulawesi water rat)
Indonesia Size: About 13 cm (5 in)[322]

Habitats: Forest and inland wetlands[477]

Diet: Plant material and invertebrates[321]
Xenuromys Tate & Archbold, 1941

One species
  • X. barbatus (Mimic tree rat)
New Guinea Size: 27–34 cm (11–13 in) long, plus 22–28 cm (9–11 in) tail[322]

Habitats: Forest and rocky areas[478]

Diet: Fruit, seeds, and insects[321]
Xeromys

Gray mouse

Thomas, 1889

One species
Papua New Guinea and northern and eastern Australia Size: 7–13 cm (3–5 in) long, plus 6–10 cm (2–4 in) tail[322]

Habitats: Intertidal marine, inland wetlands, coastal marine, and forest[479]

Diet: Insects, fish, lizards, and crabs[321]
Zelotomys
(broad-headed mouse)

Brown mouse

Osgood, 1910

Two species
  • Z. hildegardeae (Hildegarde's broad-headed mouse)
  • Z. woosnami (Woosnam's broad-headed mouse, pictured)
Central and southern Africa Size range: 9–16 cm (4–6 in) long, plus 7–14 cm (3–6 in) tail (Woosnam's broad-headed mouse)[322]

Habitats: Inland wetlands, savanna, forest, and grassland[480]

Diets: Grain, as well as insects[321]
Zyzomys
(rock rat)

Gray rat

Thomas, 1909

Five species
Australia Size range: 8 cm (3 in) long, plus 9 cm (4 in) tail (common rock rat) to 17 cm (7 in) long, plus 15 cm (6 in) tail (Arnhem Land rock rat)[322]

Habitats: Savanna, grassland, forest, rocky areas, and shrubland[481]

Diets: Fruit, seeds, other vegetation, and insects[321]
Family Nesomyidae

Members of the Nesomyidae family are called nesomyids and include Malagasy rodents, climbing mice, African rock mice, pouched rats, and tufted-tailed rats. They are generally omnivores, and eat seeds, fruit, nuts, roots, stems, and insects.[482] Nesomyidae comprises 67 extant species divided into 21 genera, grouped into six subfamilies: Cricetomyinae, containing 8 species of hamster-rats and pouched rats; Delanymyinae, containing a single species; Dendromurinae, containing 26 species of climbing mice and fat mice; Mystromyinae, containing a single species; Nesomyinae, containing 27 species of short-tailed rats, tufted-tailed rats, and big-footed mice; and Petromyscinae, containing 4 species of rock mice.

Subfamily CricetomyinaeRoberts, 1951 – three genera
Name Authority and species Range Size and ecology
Beamys
(hamster-rat)

Gray rat

Thomas, 1909

Two species
  • B. hindei (Lesser hamster-rat, pictured)
  • B. major (Greater hamster-rat)
Southeastern Africa Size range: 12 cm (5 in) long, plus 11 cm (4 in) tail (lesser hamster-rat) to 18 cm (7 in) long, plus 15 cm (6 in) tail (greater hamster-rat)[483]

Habitats: Forest[484]

Diets: Seeds and fruit[482]
Cricetomys
(pouched rat)

Gray rat

Waterhouse, 1840

Four species
Central, western, and southern Africa Size range: 28 cm (11 in) long, plus 25 cm (10 in) tail (Gambian pouched rat) to 41 cm (16 in) long, plus 45 cm (18 in) tail (southern giant pouched rat)[483]

Habitats: Forest and savanna[485]

Diets: Omnivorous, including fruit and nuts as well as vegetables, insects, crabs, and snails[482]
Saccostomus

Gray mouse

Peters, 1846

Two species
  • S. campestris (South African pouched mouse, pictured)
  • S. mearnsi (Mearns's pouched mouse)
Eastern and southern Africa Size range: 9–18 cm (4–7 in) long, plus 2–6 cm (1–2 in) tail (South African pouched mouse)[483]

Habitats: Savanna, grassland, desert, and shrubland[486]

Diets: Seeds, berries, grain, and nuts, as well as insects[482]
Subfamily DelanymyinaeMusser & Carleton, 2005 – one genus
Name Authority and species Range Size and ecology
Delanymys Hayman, 1962

One species
Central Africa
Map of range
Size: 5–7 cm (2–3 in) long, plus 8–12 cm (3–5 in) tail[483]

Habitats: Shrubland and inland wetlands[487]

Diet: Seeds[482]
Subfamily Dendromurinae – Alston, 1876 – six genera
Name Authority and species Range Size and ecology
Dendromus
(climbing mouse)

Brown mouse

Smith, 1829

Fourteen species
  • D. insignis (Montane African climbing mouse)
  • D. kahuziensis (Mount Kahuzi climbing mouse)
  • D. lachaisei (Lachaise's climbing mouse)
  • D. leucostomus (Monard's African climbing mouse)
  • D. lovati (Lovat's climbing mouse)
  • D. melanotis (Gray climbing mouse)
  • D. mesomelas (Brants's climbing mouse, pictured)
  • D. messorius (Banana climbing mouse)
  • D. mystacalis (Chestnut climbing mouse)
  • D. nyasae (Kivu climbing mouse)
  • D. nyikae (Nyika climbing mouse)
  • D. oreas (Cameroon climbing mouse)
  • D. ruppi (Rupp's African climbing mouse)
  • D. vernayi (Vernay's climbing mouse)
Sub-Saharan Africa Size range: 4 cm (2 in) long, plus 6 cm (2 in) tail (chestnut climbing mouse) to 10 cm (4 in) long, plus 9 cm (4 in) tail (Lovat's climbing mouse)[483]

Habitats: Shrubland, forest, savanna, desert, grassland, and inland wetlands[488]

Diets: Seeds, berries, insects, eggs, small lizards, and small birds[482]
Dendroprionomys Petter, 1966

One species
  • D. rousseloti (Velvet climbing mouse)
Republic of the Congo Size: About 8 cm (3 in)[483]

Habitats: Unknown[489]

Diet: Insects[482]
Malacothrix Candolle, 1838

One species
  • M. typica (Gerbil mouse)
Southern Africa Size: 6–9 cm (2–4 in) long, plus 2–5 cm (1–2 in) tail[483]

Habitats: Desert, shrubland, and savanna[490]

Diet: Green vegetation[482]
Megadendromus Dieterlen & Rupp, 1978

One species
Ethiopia Size: 11–13 cm (4–5 in) long, plus 9–11 cm (4 in) tail[483]

Habitats: Shrubland[491]

Diet: Likely vegetation and insects[482]
Prionomys

Stamp with gray mouse

Dollman, 1910

One species
West-central Africa Size: 7–9 cm (3–4 in) long, plus 10–12 cm (4–5 in) tail[483]

Habitats: Forest[492]

Diet: Insects[482]
Steatomys
(fat mouse)

Brown mouse

Peters, 1846

Eight species
  • S. bocagei (Bocage's African fat mouse)
  • S. caurinus (Northwestern fat mouse)
  • S. cuppedius (Dainty fat mouse)
  • S. jacksoni (Jackson's fat mouse)
  • S. krebsii (Krebs's fat mouse)
  • S. opimus (Pousargues African fat mouse)
  • S. parvus (Tiny fat mouse)
  • S. pratensis (Fat mouse, pictured)
Sub-Saharan Africa Size range: 5 cm (2 in) long, plus 3 cm (1 in) tail (tiny fat mouse) to 13 cm (5 in) long, plus 8 cm (3 in) tail (Bocage's African fat mouse)[483]

Habitats: Shrubland, forest, savanna, and grassland[493]

Diets: Seeds, grass bulbs, and insects[482]
Subfamily MystromyinaeVorontsov, 1966 – one genus
Name Authority and species Range Size and ecology
Mystromys Wagner, 1841

One species
Southern Africa Size: 11–20 cm (4–8 in) long, plus 4–9 cm (2–4 in) tail[483]

Habitats: Shrubland and grassland[494]

Diet: Seeds, vegetable matter, and insects[482]
Subfamily NesomyinaeForsyth Major, 1897 – nine genera
Name Authority and species Range Size and ecology
Brachytarsomys
(short-tailed rat)

Drawing of brown rodent

Günther, 1875

Two species
  • B. albicauda (White-tailed antsangy, pictured)
  • B. villosa (Hairy-tailed antsangy)
Northern and eastern Madagascar Size range: 22 cm (9 in) long, plus 22 cm (9 in) tail (white-tailed antsangy) to 24 cm (9 in) long, plus 27 cm (11 in) tail (hairy-tailed antsangy)[483]

Habitats: Forest[495]

Diets: Fruit[482]
Brachyuromys Forsyth Major, 1896

Two species
  • B. betsileoensis (Betsileo short-tailed rat)
  • B. ramirohitra (Gregarious short-tailed rat)
Eastern Madagascar Size range: 14–19 cm (6–7 in) long, plus 7–10 cm (3–4 in) tail (Betsileo short-tailed rat)[483]

Habitats: Forest, grassland, inland wetlands, and shrubland[496]

Diets: Likely vegetation and insects[482]
Eliurus
(tufted-tailed rat)

Brown rat

A. Milne-Edwards, 1885

Twelve species
  • E. antsingy (Tsingy tufted-tailed rat)
  • E. carletoni (Ankarana Special Reserve tufted-tailed rat)
  • E. danieli (Daniel's tufted-tailed rat)
  • E. ellermani (Ellerman's tufted-tailed rat)
  • E. grandidieri (Grandidier's tufted-tailed rat)
  • E. majori (Major's tufted-tailed rat)
  • E. minor (Lesser tufted-tailed rat)
  • E. myoxinus (Dormouse tufted-tailed rat, pictured)
  • E. penicillatus (White-tipped tufted-tailed rat)
  • E. petteri (Petter's tufted-tailed rat)
  • E. tanala (Tanala tufted-tailed rat)
  • E. webbi (Webb's tufted-tailed rat)
Madagascar Size range: 10 cm (4 in) long, plus 11 cm (4 in) tail (lesser tufted-tailed rat) to 17 cm (7 in) long, plus 18 cm (7 in) tail (Grandidier's tufted-tailed rat)[483]

Habitats: Forest and shrubland[497]

Diets: Likely vegetation and insects[482]
Gymnuromys Forsyth Major, 1896

One species
  • G. roberti (Voalavoanala)
Eastern Madagascar
Map of range
Size: 15–18 cm (6–7 in) long, plus 14–20 cm (6–8 in) tail[483]

Habitats: Forest[498]

Diet: Likely vegetation and insects[482]
Hypogeomys

Gray rat

Grandidier, 1869

One species
Western Madagascar
Map of range
Size: 30–34 cm (12–13 in) long, plus 21–24 cm (8–9 in) tail[483]

Habitats: Forest[499]

Diet: Fruit[482]
Macrotarsomys
(big-footed mouse)

Gray mouse

A. Milne-Edwards & Grandidier, 1898

Three species
Madagascar Size range: 8 cm (3 in) long, plus 12 cm (5 in) tail (bastard big-footed mouse) to 16 cm (6 in) long, plus 24 cm (9 in) tail (Petter's big-footed mouse)[483]

Habitats: Forest and shrubland[500]

Diets: Berries, fruit, seeds, roots, and stems[482]
Monticolomys Goodman & Carleton, 1996

One species
Madagascar
Map of range
Size: 9–11 cm (4 in) long, plus 13–15 cm (5–6 in) tail[483]

Habitats: Forest[501]

Diet: Likely vegetation and insects[482]
Nesomys
(red forest rat)

Brown mouse

Peters, 1870

Three species
  • N. audeberti (White-bellied nesomys, pictured)
  • N. lambertoni (Western nesomys)
  • N. rufus (Island mouse)
Western and eastern Madagascar Size range: 17–20 cm (7–8 in) long, plus 16–18 cm (6–7 in) tail (island mouse)[483]

Habitats: Forest[502]

Diets: Fruit and nuts[482]
Voalavo
(voalavo)
Goodman & Carleton, 1998

Two species
Central and northern Madagascar
Map of range
Size range: 8–10 cm (3–4 in) long, plus 10–13 cm (4–5 in) tail (eastern voalavo)[483]

Habitats: Forest[503]

Diets: Likely vegetation and insects[482]
Subfamily Petromyscinae – Roberts, 1951 – one genus
Name Authority and species Range Size and ecology
Petromyscus
(rock mouse)
Thomas, 1926

Four species
  • P. barbouri (Barbour's rock mouse)
  • P. collinus (Pygmy rock mouse)
  • P. monticularis (Brukkaros pygmy rock mouse)
  • P. shortridgei (Shortridge's rock mouse)
Southern Africa Size range: 6–10 cm (2–4 in) long, plus 6–9 cm (2–4 in) tail (Brukkaros pygmy rock mouse)[483]

Habitats: Rocky areas and shrubland[504]

Diets: Omnivorous[482]
Family Platacanthomyidae

Members of the Platacanthomyidae family are called platacanthomyids or Oriental dormice. They generally eat fruit, seeds, grain, and roots.[505] Platacanthomyidae comprises two extant species in two genera.

Not assigned to a named subfamily – two genera
Name Authority and species Range Size and ecology
Platacanthomys

Brown and white dormouse

Blyth, 1859

One species
Southern India
Map of range
Size: 11–14 cm (4–6 in) long, plus 7–11 cm (3–4 in) tail[506]

Habitats: Forest[507]

Diet: Fruit, seeds, grain, and roots[505]
Typhlomys

Black dormouse

Milne-Edwards, 1877

One species
Eastern China and northern Vietnam
Map of range
Size: 7–8 cm (3 in) long, plus 9–11 cm (4 in) tail[506]

Habitats: Forest[508]

Diet: Vegetation[505]
Family Spalacidae

Members of the Spalacidae family are called spalacids and include blind mole-rats, bamboo rats, mole-rats, and zokors. They are primarily herbivores, and eat roots, bulbs, tubers, grass, and seeds, with some species also eating insects.[509] Spalacidae comprises 23 extant species in 7 genera, divided into 3 subfamilies: Myospalacinae contains 6 species of zokors, Rhizomyinae contains 6 species of bamboo rats and mole-rats, Spalacinae contains 11 species of blind mole-rats.

Subfamily MyospalacinaeLilljeborg, 1866 – two genera
Name Authority and species Range Size and ecology
Eospalax
(Chinese zokor)

Gray zokor in a cage

Allen, 1938

Three species
  • E. fontanierii (Chinese zokor, pictured)
  • E. rothschildi (Rothschild's zokor)
  • E. smithii (Smith's zokor)
Central China Size range: 14 cm (6 in) long, plus 2 cm (0.8 in) tail (Rothschild's zokor) to 25 cm (10 in) long, plus 4 cm (2 in) tail (Smith's zokor)[510]

Habitats: Shrubland, grassland, and forest[511]

Diets: Roots and grains[509]
Myospalax
(zokor)

Brown zokor

Laxmann, 1769

Three species
  • M. aspalax (False zokor)
  • M. myospalax (Siberian zokor, pictured)
  • M. psilurus (Transbaikal zokor)
Central Asia Size range: 16 cm (6 in) long, plus 3 cm (1 in) tail (false zokor) to 27 cm (11 in) long, plus 5 cm (2 in) tail (Siberian zokor)[510]

Habitats: Shrubland, grassland, forest, and desert[512]

Diets: Roots and grains[509]
Subfamily RhizomyinaeWinge, 1887 – three genera
Name Authority and species Range Size and ecology
Cannomys

Brown bamboo rat

Thomas, 1915

One species
Southeastern Asia
Map of range
Size: 14–26 cm (6–10 in) long, plus 4–10 cm (2–4 in) tail[510]

Habitats: Forest[513]

Diet: Shrubs, shoots, and roots[509]
Rhizomys
(bamboo rat)

Taxidermied brown bamboo rat

Gray, 1831

Three species
Southern and southeastern Asia Size range: 22 cm (9 in) long, plus 5 cm (2 in) tail (Chinese bamboo rat) to 48 cm (19 in) long, plus 20 cm (8 in) tail (large bamboo rat)[510]

Habitats: Grassland and forest[514]

Diets: Bamboo roots, as well as grass, seeds, and fruit[509]
Tachyoryctes
(African mole-rat)

Brown mole-rat

Rüppell, 1835

Two species
Eastern Africa Size range: 15 cm (6 in) long, plus 4 cm (2 in) tail (northeast African mole-rat) to 31 cm (12 in) long, plus 7 cm (3 in) tail (giant root-rat)[510]

Habitats: Shrubland, grassland, savanna, and forest[515]

Diets: Roots, rhizomes, tubers, bulbs, and corms, as well as grass and legumes[509]
Subfamily SpalacinaeGray, 1821 – two genera
Name Authority and species Range Size and ecology
Nannospalax
(small-bodied mole-rat)

Gray mole-rat

Palmer, 1903

Three species
Southeastern Europe, Western Asia, and northeastern Africa Size range: 13 cm (5 in) long, plus 0 cm (0 in) tail (Middle East blind mole-rat) to 25 cm (10 in) long (Anatolian blind mole-rat)[510]

Habitats: Shrubland, grassland, and desert[516]

Diets: Roots, tubers, acorns, plant stems, and other plant parts[509]
Spalax
(blind mole-rat)

Gray mole-rat

Güldenstädt, 1770

Eight species
Eastern Europe, Kazakhstan, and southwestern Russia Size range: 19 cm (7 in) long, plus 0 cm (0 in) tail (greater blind mole-rat) to 35 cm (14 in) long (giant blind mole-rat)[510]

Habitats: Shrubland, grassland, and forest[517]

Diets: Roots, bulbs, tubers, grass, and seeds, as well as insects[509]

Suborder Sciuromorpha

Family Aplodontiidae

Members of the Aplodontiidae family are called aplodontiids. It includes a single herbivorous species.[518]

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Aplodontia

Gray beaver

Richardson, 1829

One species
Western United States and southwestern Canada Size: 23–43 cm (9–17 in) long, plus 2–6 cm (1–2 in) tail[519]

Habitats: Forest, shrubland, and inland wetlands[520]

Diet: A wide variety of vegetation[518]

Family Gliridae

Members of the Gliridae family are called glirids or dormice. They are omnivores and feed on fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents.[521] Gliridae comprises 29 extant species in 9 genera, divided into 3 subfamilies: Glirinae, containing two species; Graphiurinae, containing fifteen species; and Leithiinae, containing twelve species.

Subfamily Glirinae – Muirhead, 1819 – two genera
Name Authority and species Range Size and ecology
Glirulus

Brown and white dormice

Thomas, 1906

One species
Japan
Map of range
Size: 6–10 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[522]

Habitats: Forest[523]

Diet: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents[521]
Glis

Brown and white dormouse

Brisson, 1762

One species
Europe and western Asia
Map of range
Size: 13–19 cm (5–7 in) long, plus 10–18 cm (4–7 in) tail[522]

Habitats: Forest and shrubland[524]

Diet: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents[521]
Subfamily GraphiurinaeWinge, 1887 – one genus
Name Authority and species Range Size and ecology
Graphiurus
(African dormouse)

Brown and white dormouse

Smuts, 1832

Fifteen species
  • G. angolensis (Angolan African dormouse)
  • G. christyi (Christy's dormouse)
  • G. crassicaudatus (Jentink's dormouse)
  • G. johnstoni (Johnston's African dormouse)
  • G. kelleni (Kellen's dormouse)
  • G. lorraineus (Lorrain dormouse)
  • G. microtis (Small-eared dormouse)
  • G. monardi (Monard's dormouse)
  • G. murinus (Woodland dormouse, pictured)
  • G. nagtglasii (Nagtglas's African dormouse)
  • G. ocularis (Spectacled dormouse)
  • G. platyops (Rock dormouse)
  • G. rupicola (Stone dormouse)
  • G. surdus (Silent dormouse)
  • G. walterverheyeni (Walter Verheyen's African dormouse)
Sub-Saharan Africa Size range: 6 cm (2 in) long, plus 6 cm (2 in) tail (Johnston's African dormouse) to 16 cm (6 in) long, plus 13 cm (5 in) tail (Monard's dormouse)[522]

Habitats: Grassland, inland wetlands, rocky areas, savanna, shrubland, and forest[525]

Diets: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents[521]
Subfamily Leithiinae – Lydekker, 1896 – six genera
Name Authority and species Range Size and ecology
Chaetocauda Wang, 1985

One species
  • C. sichuanensis (Chinese dormouse)
Central China Size: 9–10 cm (4 in) long, plus 9–11 cm (4 in) tail[522]

Habitats: Forest[526]

Diet: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents[521]
Dryomys
(forest dormouse)

Brown and white dormouse

Thomas, 1905

Three species
  • D. laniger (Woolly dormouse)
  • D. niethammeri (Balochistan forest dormouse)
  • D. nitedula (Forest dormouse, pictured)
Eastern Europe and western and central Asia Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (forest dormouse) to 11 cm (4 in) long, plus 9 cm (4 in) tail (Balochistan forest dormouse)[522]

Habitats: Shrubland, forest, and rocky areas[527]

Diets: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents[521]
Eliomys
(garden dormouse)

Brown and white dormouse

Wagner, 1840

Three species
Europe, western Asia, and northern Africa
Map of range
Size range: 9 cm (4 in) long, plus 8 cm (3 in) tail (garden dormouse) to 15 cm (6 in) long, plus 14 cm (6 in) tail (Asian garden dormouse)[522]

Habitats: Desert, rocky areas, coastal marine, shrubland, and forest[528]

Diets: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents[521]
Muscardinus

Brown dormouse

Kaup, 1829

One species
Europe and western Asia
Map of range
Size: 6–10 cm (2–4 in) long, plus 5–9 cm (2–4 in) tail[522]

Habitats: Forest[529]

Diet: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents[521]
Myomimus
(mouse-tailed dormouse)
Ogniov, 1924

Three species
Western Asia and southeastern Europe Size range: 6 cm (2 in) long, plus 6 cm (2 in) tail (Setzer's mouse-tailed dormouse) to 14 cm (6 in) long, plus 10 cm (4 in) tail (Roach's mouse-tailed dormouse)[522]

Habitats: Shrubland, forest, and savanna[530]

Diets: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents[521]
Selevinia

Brown and white dormouse on stamp

Belosludov & Bazhanov, 1939

One species
Kazakhstan Size: 7–10 cm (3–4 in) long, plus 5–8 cm (2–3 in) tail[522]

Habitats: Desert[531]

Diet: Insects and spiders[521]

Family Sciuridae

Members of the Sciuridae family are called sciurids and include squirrels, chipmunks, and marmots. They primarily feed on fruit, seeds, and nuts, though many species also eat insects, fungi, bark, eggs, and small vertebrates.[532] Sciuridae comprises 284 extant species in 60 genera, divided into 5 subfamilies: Callosciurinae, containing 67 species of Asian squirrels; Ratufinae, containing 4 species of giant squirrels; Sciurillinae, containing a single species of pygmy squirrel; Sciurinae, containing 84 species of flying squirrel and tree squirrels; and Xerinae, containing 128 species of marmots, chipmunks, prairie dogs, and ground squirrels.

Subfamily CallosciurinaePocock, 1923 – fourteen genera
Name Authority and species Range Size and ecology
Callosciurus
(beautiful squirrel)

Gray squirrel

Gray, 1867

Fifteen species
  • C. adamsi (Ear-spot squirrel)
  • C. albescens (Kloss's squirrel)
  • C. baluensis (Kinabalu squirrel)
  • C. caniceps (Grey-bellied squirrel)
  • C. erythraeus (Pallas's squirrel)
  • C. finlaysonii (Finlayson's squirrel)
  • C. inornatus (Inornate squirrel)
  • C. melanogaster (Mentawai squirrel)
  • C. nigrovittatus (Black-striped squirrel, pictured)
  • C. notatus (Plantain squirrel)
  • C. orestes (Borneo black-banded squirrel)
  • C. phayrei (Phayre's squirrel)
  • C. prevostii (Prevost's squirrel)
  • C. pygerythrus (Irrawaddy squirrel)
  • C. quinquestriatus (Anderson's squirrel)
Southeastern and eastern Asia Size range: 14 cm (6 in) long, plus 13 cm (5 in) tail (Borneo black-banded squirrel) to 29 cm (11 in) long, plus 21 cm (8 in) tail (Inornate squirrel)[533]

Habitats: Shrubland, inland wetlands, and forest[534]

Diets: Fruit, nuts, seeds, flowers, vegetation, insects, and eggs[532]
Dremomys
(red-cheeked squirrel)

Brown squirrel

Heude, 1898

Six species
  • D. everetti (Bornean mountain ground squirrel)
  • D. gularis (Red-throated squirrel)
  • D. lokriah (Orange-bellied Himalayan squirrel, pictured)
  • D. pernyi (Perny's long-nosed squirrel)
  • D. pyrrhomerus (Red-hipped squirrel)
  • D. rufigenis (Asian red-cheeked squirrel)
Southern, southeastern, and eastern Asia Size range: 15 cm (6 in) long, plus 9 cm (4 in) tail (Bornean mountain ground squirrel) to 22 cm (9 in) long, plus 17 cm (7 in) tail (red-throated squirrel)[533]

Habitats: Shrubland, rocky areas, and forest[535]

Diets: Fruit, nuts, vegetation, and insects[532]
Exilisciurus
(pygmy squirrel)

Brown squirrel

Moore, 1958

Three species
  • E. concinnus (Philippine pygmy squirrel)
  • E. exilis (Least pygmy squirrel)
  • E. whiteheadi (Tufted pygmy squirrel, pictured)
Southeastern Asia Size range: 7 cm (3 in) long, plus 4 cm (2 in) tail (least pygmy squirrel) to 9 cm (4 in) long, plus 7 cm (3 in) tail (Philippine pygmy squirrel)[533]

Habitats: Forest[536]

Diets: Vegetation and insects[532]
Funambulus
(palm squirrel)

Gray and white squirrel

Lesson, 1835

Six species
  • F. layardi (Layard's palm squirrel)
  • F. obscurus (Dusky striped squirrel)
  • F. palmarum (Indian palm squirrel, pictured)
  • F. pennantii (Northern palm squirrel)
  • F. sublineatus (Nilgiri striped squirrel)
  • F. tristriatus (Jungle palm squirrel)
Southern Asia Size range: 10 cm (4 in) long, plus 7 cm (3 in) tail (dusky striped squirrel) to 16 cm (6 in) long, plus 15 cm (6 in) tail (jungle palm squirrel)[533]

Habitats: Shrubland, grassland, inland wetlands, and forest[537]

Diets: Seeds, nuts, bark, buds, leaves, flowers, and insects[532]
Glyphotes

Brown squirrel

Thomas, 1898

One species
  • G. simus (Sculptor squirrel)
Borneo Size: 9–15 cm (4–6 in) long, plus 9–11 cm (4 in) tail[533]

Habitats: Forest[538]

Diet: Fruit, nuts, seeds, flowers, vegetation, insects, and eggs[532]
Hyosciurus Archbold & Tate, 1935

Two species
  • H. heinrichi (Montane long-nosed squirrel)
  • H. ileile (Lowland long-nosed squirrel)
Indonesia Size range: 19 cm (7 in) long, plus 6 cm (2 in) tail (Montane long-nosed squirrel) to 25 cm (10 in) long, plus 13 cm (5 in) tail (Lowland long-nosed squirrel)[533]

Habitats: Shrubland and forest[539]

Diets: Fruit, nuts, and insects[532]
Lariscus

Drawing of brown squirrel

Thomas & Wroughton, 1909

Four species
  • L. hosei (Four-striped ground squirrel)
  • L. insignis (Three-striped ground squirrel, pictured)
  • L. niobe (Niobe ground squirrel)
  • L. obscurus (Mentawai three-striped squirrel)
Southeastern Asia Size range: 18 cm (7 in) long, plus 8 cm (3 in) tail (Niobe ground squirrel) to 20 cm (8 in) long, plus 9 cm (4 in) tail (Mentawai three-striped squirrel)[533]

Habitats: Shrubland and forest[540]

Diets: Fruit and nuts[532]
Menetes

Brown squirrel

Thomas, 1908

One species
  • M. berdmorei (Berdmore's ground squirrel)
Southeastern Asia
Map of range
Size: About 18 cm (7 in)[533]

Habitats: Grassland, shrubland, and forest[541]

Diet: Fruit and nuts[532]
Nannosciurus

Brown squirrels

Trouessart, 1880

One species
  • N. melanotis (Black-eared squirrel)
Southeastern Asia Size: About 8 cm (3 in)[533]

Habitats: Forest[542]

Diet: Fruit and nuts[532]
Prosciurillus

Gray squirrel

Ellerman, 1947

Seven species
  • P. abstrusus (Secretive dwarf squirrel)
  • P. alstoni (Alston's squirrel)
  • P. leucomus (Whitish dwarf squirrel)
  • P. murinus (Celebes dwarf squirrel)
  • P. rosenbergii (Sanghir squirrel, pictured)
  • P. topapuensis (Mount Topapu squirrel)
  • P. weberi (Weber's dwarf squirrel)
Indonesia and Philippines Size range: 10 cm (4 in) long, plus 5 cm (2 in) tail (Celebes dwarf squirrel) to 20 cm (8 in) long, plus 18 cm (7 in) tail (Alston's squirrel)[533]

Habitats: Forest[543]

Diets: Fruit, nuts, bark, and resin[532]
Rhinosciurus

Brown squirrel

Blyth, 1856

One species
  • R. laticaudatus (Shrew-faced squirrel)
Southeastern Asia
Map of range
Size: 19–21 cm (7–8 in) long, plus 11–14 cm (4–6 in) tail[533]

Habitats: Forest[544]

Diet: Insects and earthworms, as well as fruit[532]
Rubrisciurus Ellerman, 1954

One species
Island of Sulawesi in Indonesia Size: 15–30 cm (6–12 in) long, plus 18–25 cm (7–10 in) tail[533]

Habitats: Forest[545]

Diet: Fruit and seeds[532]
Sundasciurus
(Sunda squirrel)

Gray squirrel

Moore, 1958

Fifteen species
  • S. brookei (Brooke's squirrel)
  • S. davensis (Davao squirrel)
  • S. fraterculus (Fraternal squirrel)
  • S. hippurus (Horse-tailed squirrel)
  • S. hoogstraali (Busuanga squirrel)
  • S. jentinki (Jentink's squirrel)
  • S. juvencus (Northern Palawan tree squirrel)
  • S. lowii (Low's squirrel)
  • S. mindanensis (Mindanao squirrel)
  • S. moellendorffi (Culion tree squirrel)
  • S. philippinensis (Philippine tree squirrel)
  • S. rabori (Palawan montane squirrel)
  • S. samarensis (Samar squirrel)
  • S. steerii (Southern Palawan tree squirrel)
  • S. tenuis (Slender squirrel, pictured)
Southeastern Asia Size range: 11 cm (4 in) long, plus 7 cm (3 in) tail (fraternal squirrel) to 25 cm (10 in) long, plus 25 cm (10 in) tail (horse-tailed squirrel)[533]

Habitats: Forest[546]

Diets: Fruit and vegetation, as well as insects[532]
Tamiops
(Asiatic striped squirrel)

Brown squirrel

Allen, 1906

Four species
  • T. maritimus (Maritime striped squirrel)
  • T. mcclellandii (Himalayan striped squirrel, pictured)
  • T. rodolphii (Cambodian striped squirrel)
  • T. swinhoei (Swinhoe's striped squirrel)
Southeastern and eastern Asia Size range: 11 cm (4 in) long, plus 10 cm (4 in) tail (Cambodian striped squirrel) to 13 cm (5 in) long, plus 11 cm (4 in) tail (maritime striped squirrel)[533]

Habitats: Shrubland, grassland, inland wetlands, and forest[547]

Diets: Fruit, nuts, seeds, and insects[532]
Subfamily RatufinaeMoore, 1959 – one genus
Name Authority and species Range Size and ecology
Ratufa
(giant squirrel)

Gray squirrel

Gray, 1867

Four species
Southern and southeastern Asia Size range: 32 cm (13 in) long, plus 35 cm (14 in) tail (grizzled giant squirrel) to 45 cm (18 in) long, plus 49 cm (19 in) tail (Indian giant squirrel)[533]

Habitats: Forest[548]

Diets: Fruit, nuts, bark, insects, and eggs[532]
Subfamily SciurillinaeMoore, 1959 – one genus
Name Authority and species Range Size and ecology
Sciurillus

Drawing of brown squirrel

Thomas, 1914

One species
Northern South America
Map of range
Size: 8–12 cm (3–5 in) long, plus 8–12 cm (3–5 in) tail[533]

Habitats: Forest[549]

Diet: Fruit, nuts, bark, and resin[532]
Subfamily SciurinaeHemprich, 1820 – 20 genera
Name Authority and species Range Size and ecology
Aeretes

Drawing of brown squirrel

Allen, 1940

One species
  • A. melanopterus (Groove-toothed flying squirrel)
Central China Size: 27–35 cm (11–14 in) long, plus 27–36 cm (11–14 in) tail[533]

Habitats: Forest[550]

Diet: Nuts, fruit, twigs, shoots, leaves, bark and insects[532]
Aeromys
(large black flying squirrel)

Drawing of black squirrel

Robinson & Kloss, 1915

Two species
  • A. tephromelas (Black flying squirrel, pictured)
  • A. thomasi (Thomas's flying squirrel)
Southeastern Asia and Indonesia and Malaysia Size range: 30 cm (12 in) long, plus 37 cm (15 in) tail (Thomas's flying squirrel) to 38 cm (15 in) long, plus 44 cm (17 in) tail (black flying squirrel)[533]

Habitats: Forest[551]

Diets: Fruit, nuts, leaves, and insects[532]
Belomys

Drawing of brown squirrels

Thomas, 1908

One species
  • B. pearsonii (Hairy-footed flying squirrel)
Southeastern and eastern Asia
Map of range
Size: 18–21 cm (7–8 in) long, plus 13–17 cm (5–7 in) tail[533]

Habitats: Forest[552]

Diet: Nuts, fruit, twigs, shoots, leaves, bark and insects[532]
Biswamoyopterus
(Indochinese giant flying squirrel)
Saha, 1981

Two species
Laos and eastern India Size range: 40 cm (16 in) long, plus 60 cm (24 in) tail (Namdapha flying squirrel) to 46 cm (18 in) long, plus 62 cm (24 in) tail (Laotian giant flying squirrel)[533]

Habitats: Forest[553]

Diets: Unknown[532]
Eoglaucomys Howell, 1915

One species
Central Asia Size: 23–30 cm (9–12 in) long, plus 25–33 cm (10–13 in) tail[533]

Habitats: Forest[554]

Diet: Seeds, as well as shoots, buds, leaves, and nuts[532]
Eupetaurus

Drawing of brown squirrel

Thomas, 1888

One species
  • E. cinereus (Western woolly flying squirrel)
Western China Size: 42–61 cm (17–24 in) long, plus 38–54 cm (15–21 in) tail[533]

Habitats: Forest, rocky areas, and caves[555]

Diet: Moss and lichen[532]
Glaucomys
(New World flying squirrel)

Brown squirrel

Thomas, 1908

Two species
Eastern North America and Canada and northern United States Size range: 11–19 cm (4–7 in) long, plus 12–16 cm (5–6 in) tail (Northern flying squirrel)[533]

Habitats: Forest[556]

Diets: Nuts, bark, lichen, fungi, fruit, and berries, as well as insects[532]
Hylopetes
(arrow-tailed flying squirrel)

Gray squirrel

Thomas, 1908

Nine species
  • H. alboniger (Particolored flying squirrel, pictured)
  • H. bartelsi (Bartel's flying squirrel)
  • H. nigripes (Palawan flying squirrel)
  • H. phayrei (Indochinese flying squirrel)
  • H. platyurus (Jentink's flying squirrel)
  • H. sagitta (Arrow flying squirrel)
  • H. sipora (Sipora flying squirrel)
  • H. spadiceus (Red-cheeked flying squirrel)
  • H. winstoni (Sumatran flying squirrel)
Southeastern and eastern Asia Size range: 11 cm (4 in) long, plus 9 cm (4 in) tail (arrow flying squirrel) to 33 cm (13 in) long, plus 31 cm (12 in) tail (Palawan flying squirrel)[533]

Habitats: Shrubland and forest[557]

Diets: Nuts, shoots, leaves, fruit, insects, and small snakes[532]
Iomys
(Horsfield's flying squirrel)

Drawing of brown squirrel

Thomas, 1908

Two species
  • I. horsfieldii (Javanese flying squirrel, pictured)
  • I. sipora (Mentawi flying squirrel)
Southeastern Asia and Indonesia Size range: 16–23 cm (6–9 in) long, plus 15–23 cm (6–9 in) tail (Javanese flying squirrel)[533]

Habitats: Shrubland and forest[558]

Diets: Nuts, fruit, twigs, shoots, leaves, bark and insects[532]
Microsciurus
(dwarf squirrel)

Gray squirrel

Allen, 1895

Four species
Central America and northern South America Size range: 10 cm (4 in) long, plus 8 cm (3 in) tail (Central American dwarf squirrel) to 16 cm (6 in) long, plus 15 cm (6 in) tail (Amazon dwarf squirrel)[533]

Habitats: Forest[559]

Diets: Fruit and nuts[532]
Petaurillus
(pygmy flying squirrel)
Thomas, 1908

Three species
  • P. emiliae (Lesser pygmy flying squirrel)
  • P. hosei (Hose's pygmy flying squirrel)
  • P. kinlochii (Selangor pygmy flying squirrel)
Borneo and Malaysia Size range: 6 cm (2 in) long, plus 6 cm (2 in) tail (lesser pygmy flying squirrel) to 10 cm (4 in) long, plus 10 cm (4 in) tail (Selangor pygmy flying squirrel)[533]

Habitats: Forest[560]

Diets: Nuts, fruit, twigs, shoots, leaves, bark and insects[532]
Petaurista
(giant flying squirrel)

Brown and white squirrel

Link, 1795

Ten species
Asia Size range: 27 cm (11 in) long, plus 28 cm (11 in) tail (Japanese giant flying squirrel) to 61 cm (24 in) long, plus 69 cm (27 in) tail (Indian giant flying squirrel)[533]

Habitats: Rocky areas and forest[561]

Diets: Fruit, nuts, twigs, shoots, and leaves, as well as insects[532]
Petinomys
(dwarf flying squirrel)

Drawing of squirrel

Thomas, 1908

Eight species
  • P. crinitus (Basilan flying squirrel)
  • P. fuscocapillus (Travancore flying squirrel)
  • P. genibarbis (Whiskered flying squirrel, pictured)
  • P. hageni (Hagen's flying squirrel)
  • P. lugens (Siberut flying squirrel)
  • P. mindanensis (Mindanao flying squirrel)
  • P. setosus (Temminck's flying squirrel)
  • P. vordermanni (Vordermann's flying squirrel)
Southeastern Asia and southern India Size range: 9 cm (4 in) long, plus 8 cm (3 in) tail (Temminck's flying squirrel) to 37 cm (15 in) long, plus 46 cm (18 in) tail (Mindanao flying squirrel)[533]

Habitats: Forest[562]

Diets: Nuts, fruit, twigs, shoots, and leaves, as well as bark and insects[532]
Pteromys
(Old World flying squirrel)

Brown squirrel

Cuvier, 1800

Two species
Japan, northern Asia, and northern Europe Size range: 12–23 cm (5–9 in) long, plus 9–15 cm (4–6 in) tail (Siberian flying squirrel)[533]

Habitats: Forest[563]

Diets: Nuts, pine seeds, buds, bark, fruit, and insects[532]
Pteromyscus

Drawing of gray squirrel

Thomas, 1908

One species
  • P. pulverulentus (Smoky flying squirrel)
Southeastern Asia Size: 20–29 cm (8–11 in) long, plus 17–23 cm (7–9 in) tail[533]

Habitats: Forest[564]

Diet: Nuts, fruit, twigs, shoots, leaves, bark and insects[532]
Rheithrosciurus

Brown, black, and white squirrel

Gray, 1867

One species
Borneo
Map of range
Size: 33–35 cm (13–14 in) long, plus 30–34 cm (12–13 in) tail[533]

Habitats: Forest[565]

Diet: Fruit and nuts[532]
Sciurus
(tree squirrel)

Brown squirrel

Linnaeus, 1758

29 species
North America, South America, Europe, and Asia Size range: 14 cm (6 in) long, plus 11 cm (4 in) tail (Andean squirrel) to 56 cm (22 in) long, plus 27 cm (11 in) tail (Peters's squirrel)[533]

Habitats: Shrubland, grassland, and forest[566]

Diets: Nuts, seeds, fruit, buds, and tree shoots[532]
Syntheosciurus

Brown squirrel

Bangs, 1902

One species
Costa Rica and Panama Size: 15–19 cm (6–7 in) long, plus 12–16 cm (5–6 in) tail[533]

Habitats: Forest[567]

Diet: Flowers, buds, leaves, and sap[532]
Tamiasciurus
(American pine squirrel)

Brown squirrel

Trouessart, 1880

Three species
North America Size range: 18–19 cm (7 in) long, plus 11–15 cm (4–6 in) tail (multiple)[533]

Habitats: Forest[568]

Diets: Nuts, buds, fruit, bark, fungi, sap, eggs, and small vertebrates[532]
Trogopterus

Drawing of brown squirrel

Heude, 1898

One species
Southern China Size: 20–33 cm (8–13 in) long, plus 26–30 cm (10–12 in) tail[533]

Habitats: Forest and caves[569]

Diet: Leaves, twigs, fruit, and nuts[532]
Subfamily XerinaeOsborn, 1910 – 24 genera
Name Authority and species Range Size and ecology
Ammospermophilus
(antelope squirrel)

Brown squirrel

Merriam, 1892

Four species
Western United States and western Mexico Size range: 18 cm (7 in) long, plus 4 cm (2 in) tail (white-tailed antelope squirrel) to 28 cm (11 in) long, plus 10 cm (4 in) tail (Harris's antelope squirrel)[533]

Habitats: Shrubland, grassland, desert, and forest[570]

Diets: Seeds, fruit, stems, and roots, as well as insects and carrion[532]
Atlantoxerus

Brown and white squirrel

Forsyth Major, 1893

One species
Northwestern Africa
Map of range
Size: About 17 cm (7 in)[533]

Habitats: Rocky areas, grassland, and shrubland[571]

Diet: Fruit and seeds[532]
Callospermophilus
(golden-mantled ground squirrel)

Brown squirrel

Merriam, 1897

Three species
Western North America Size range: 16 cm (6 in) long, plus 6 cm (2 in) tail (Sierra Madre ground squirrel) to 31 cm (12 in) long, plus 12 cm (5 in) tail (Cascade golden-mantled ground squirrel)[533]

Habitats: Shrubland, rocky areas, grassland, and forest[572]

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs[532]
Cynomys
(prairie dog)

Brown prairie dog

Rafinesque, 1817

Five species
Southern Canada, western United States, and northern Mexico Size range: 29 cm (11 in) long, plus 4 cm (2 in) tail (Utah prairie dog) to 44 cm (17 in) long, plus 11 cm (4 in) tail (Mexican prairie dog)[533]

Habitats: Shrubland, savanna, grassland, and desert[573]

Diets: Herbs and grasses[532]
Epixerus

Brown squirrel

Thomas, 1909

One species
  • E. ebii (Ebian's palm squirrel)
Western Africa
Map of range
Size: 28–29 cm (11–11 in) long, plus about 28 cm (11 in) tail[533]

Habitats: Forest[574]

Diet: Fruit as well as insects[532]
Eutamias

Brown chipmunk

Trouessart, 1880

One species
Northern and eastern Asia and Europe
Map of range
Size: 14–15 cm (6 in) long, plus 10–12 cm (4–5 in) tail[533]

Habitats: Forest and shrubland[575]

Diet: Nuts and seeds, as well as buds, berries, grains, and fungi[532]
Funisciurus
(rope squirrel)

Brown squirrel

Trouessart, 1880

Ten species
  • F. anerythrus (Thomas's rope squirrel)
  • F. bayonii (Lunda rope squirrel)
  • F. carruthersi (Carruther's mountain squirrel)
  • F. congicus (Congo rope squirrel, pictured)
  • F. duchaillui (Du Chaillu's rope squirrel)
  • F. isabella (Lady Burton's rope squirrel)
  • F. lemniscatus (Ribboned rope squirrel)
  • F. leucogenys (Red-cheeked rope squirrel)
  • F. pyrropus (Fire-footed rope squirrel)
  • F. substriatus (Kintampo rope squirrel)
Central and western Africa Size range: 16 cm (6 in) long, plus 16 cm (6 in) tail (Congo rope squirrel) to 25 cm (10 in) long, plus 20 cm (8 in) tail (Lunda rope squirrel)[533]

Habitats: Shrubland, rocky areas, savanna, and forest[576]

Diets: Seeds, nuts, and fruit, as well as insects and eggs[532]
Heliosciurus
(sun squirrel)

Gray squirrel

Trouessart, 1880

Six species
Sub-Saharan Africa Size range: 18 cm (7 in) long, plus 20 cm (8 in) tail (small sun squirrel) to 24 cm (9 in) long, plus 27 cm (11 in) tail (Zanj sun squirrel)[533]

Habitats: Savanna and forest[577]

Diets: Seeds, nuts, fruit, insects, and eggs[532]
Ictidomys
(lined ground squirrel)

Brown squirrel

Allen, 1877

Two species
North America Size range: 17 cm (7 in) long, plus 6 cm (2 in) tail (thirteen-lined ground squirrel) to 38 cm (15 in) long, plus 17 cm (7 in) tail (Mexican ground squirrel)[533]

Habitats: Shrubland, grassland, and coastal marine[578]

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs[532]
Marmota
(marmot)

Brown groundhog

Blumenbach, 1779

Fourteen species
Asia, Europe, United States, and Canada Size range: 36 cm (14 in) long, plus 11 cm (4 in) tail (Tarbagan marmot) to 75 cm (30 in) long, plus 24 cm (9 in) tail (Olympic marmot)[533]

Habitats: Rocky areas, forest, grassland, shrubland, and desert[579]

Diets: Grass and forbs, as well as fruit, grains, legumes, and insects[532]
Myosciurus Thomas, 1909

One species
Western Africa
Map of range
Size: 7–8 cm (3 in) long, plus 5–6 cm (2 in) tail[533]

Habitats: Forest[580]

Diet: Fruit, bark, and insects[532]
Neotamias
(chipmunk)

Brown chipmunk

Howell, 1929

23 species
North America Size range: 10 cm (4 in) long, plus 6 cm (2 in) tail (alpine chipmunk) to 16 cm (6 in) long, plus 11 cm (4 in) tail (Durango chipmunk)[533]

Habitats: Rocky areas, forest, grassland, shrubland, and desert[581]

Diets: Fruit and seeds, as well as flowers, fungi, insects, and eggs[532]
Notocitellus
(tropical ground squirrel)

Gray squirrel

Howell, 1938

Two species
Mexico Size range: 16 cm (6 in) long, plus 13 cm (5 in) tail (tropical ground squirrel) to 22 cm (9 in) long, plus 21 cm (8 in) tail (ring-tailed ground squirrel)[533]

Habitats: Shrubland, rocky areas, and forest[582]

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs[532]
Otospermophilus
(American rock squirrel)

Gray squirrel

Brandt, 1844

Two species
Mexico and western United States Size range: 28 cm (11 in) long, plus 13 cm (5 in) tail (California ground squirrel) to 54 cm (21 in) long, plus 26 cm (10 in) tail (rock squirrel)[533]

Habitats: Shrubland, rocky areas, and grassland[583]

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs[532]
Paraxerus
(African bush squirrel)

Gray squirrel

Forsyth Major, 1893

Eleven species
  • P. alexandri (Alexander's bush squirrel)
  • P. boehmi (Boehm's bush squirrel)
  • P. cepapi (Smith's bush squirrel, pictured)
  • P. cooperi (Cooper's mountain squirrel)
  • P. flavovittis (Striped bush squirrel)
  • P. lucifer (Black and red bush squirrel)
  • P. ochraceus (Ochre bush squirrel)
  • P. palliatus (Red bush squirrel)
  • P. poensis (Green bush squirrel)
  • P. vexillarius (Swynnerton's bush squirrel)
  • P. vincenti (Vincent's bush squirrel)
Sub-Saharan Africa Size range: 13 cm (5 in) long, plus 10 cm (4 in) tail (Alexander's bush squirrel) to 23 cm (9 in) long, plus 20 cm (8 in) tail (black and red bush squirrel)[533]

Habitats: Shrubland, savanna, and forest[584]

Diets: Seeds and fruit, as well as roots and eggs[532]
Poliocitellus

Gray squirrel

Howell, 1938

One species
Central United States and southern Canada Size: 23–24 cm (9 in) long, plus about 13 cm (5 in) tail[533]

Habitats: Savanna, grassland, and inland wetlands[585]

Diet: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs[532]
Protoxerus
(African giant squirrel)

Gray squirrel

Forsyth Major, 1893

Two species
Western and central Africa Size range: 24 cm (9 in) long, plus 30 cm (12 in) tail (slender-tailed squirrel) to 30 cm (12 in) long, plus 31 cm (12 in) tail (forest giant squirrel)[533]

Habitats: Forest[586]

Diets: Seeds, nuts, and fruit[532]
Sciurotamias
(Chinese rock squirrel)

Gray squirrel

Miller, 1901

Two species
  • S. davidianus (Père David's rock squirrel, pictured)
  • S. forresti (Forrest's rock squirrel)
China Size range: 20 cm (8 in) long, plus 14 cm (6 in) tail (Père David's rock squirrel) to 22 cm (9 in) long, plus 16 cm (6 in) tail (Forrest's rock squirrel)[533]

Habitats: Shrubland and rocky areas[587]

Diets: Fruit and seeds[532]
Spermophilopsis

Brown squirrel

Blasius, 1884

One species
Central Asia Size: 23–27 cm (9–11 in) long, plus 2–9 cm (1–4 in) tail[533]

Habitats: Desert, grassland, and shrubland[588]

Diet: Fruit, seeds, vegetation, and insects[532]
Spermophilus
(Holarctic ground squirrel)

Brown squirrel

F. Cuvier, 1825

Fifteen species
Asia and eastern Europe Size range: 14 cm (6 in) long, plus 3 cm (1 in) tail (Asia Minor ground squirrel) to 32 cm (13 in) long, plus 11 cm (4 in) tail (Russet ground squirrel)[533]

Habitats: Rocky areas, grassland, shrubland, and desert[589]

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs[532]
Tamias

Brown chipmunk

Illiger, 1811

One species
Eastern United States and eastern Canada
Map of range
Size: 14–15 cm (6 in) long, plus 8–10 cm (3–4 in) tail[533]

Habitats: Shrubland and forest[590]

Diet: Nuts, seeds, fruit, and fungi[532]
Urocitellus
(ground squirrel)

Brown squirrel

Obolenskij, 1927

Twelve species
Canada, United States, and central and eastern Asia Size range: 15 cm (6 in) long, plus 3 cm (1 in) tail (Merriam's ground squirrel) to 34 cm (13 in) long, plus 9 cm (4 in) tail (Richardson's ground squirrel)[533]

Habitats: Shrubland, grassland, desert, and forest[591]

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs[532]
Xerospermophilus
(desert ground squirrel)

Brown squirrel

Merriam, 1892

Four species
Mexico and United States Size range: 18 cm (7 in) long, plus 5 cm (2 in) tail (spotted ground squirrel) to 28 cm (11 in) long, plus 12 cm (5 in) tail (round-tailed ground squirrel)[533]

Habitats: Grassland, desert, and forest[592]

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs[532]
Xerus
(unstriped ground squirrel)

Brown squirrel

Hemprich & Ehrenberg, 1833

Four species
Sub-Saharan Africa Size range: 22–29 cm (9–11 in) long, plus 18–26 cm (7–10 in) tail (African striped ground squirrel)[533]

Habitats: Shrubland, desert, savanna, and forest[593]

Diets: Roots, seeds, fruit, grains, insects, small vertebrates, and eggs[532]

References

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  359. ^ "Brassomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
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  369. ^ "Conilurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
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  380. ^ "Echiothrix". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
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  383. ^ "Golunda". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
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  397. ^ "Komodomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  398. ^ "Lamottemys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  399. ^ "Leggadina". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
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  489. ^ "Dendroprionomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  490. ^ "Malacothrix". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  491. ^ "Megadendromus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  492. ^ "Prionomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  493. ^ "Steatomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  494. ^ "Mystromys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  495. ^ "Brachytarsomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  496. ^ "Brachyuromys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  497. ^ "Eliurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  498. ^ "Gymnuromys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  499. ^ "Hypogeomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  500. ^ "Macrotarsomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  501. ^ "Monticolomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  502. ^ "Nesomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  503. ^ "Voalavo". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  504. ^ "Petromyscus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  505. ^ a b c Nowak, p. 1432
  506. ^ a b Chernasky; Motis; Burgin, p. 210
  507. ^ "Platacanthomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  508. ^ "Typhlomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  509. ^ a b c d e f g h Nowak, pp. 1427–1445
  510. ^ a b c d e f g Chernasky; Motis; Burgin, pp. 210–213
  511. ^ "Eospalax". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  512. ^ "Myospalax". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  513. ^ "Cannomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  514. ^ "Rhizomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  515. ^ "Tachyoryctes". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  516. ^ "Nannospalax". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  517. ^ "Spalax". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  518. ^ a b Nowak, p. 1245
  519. ^ Chernasky; Motis; Burgin, p. 374
  520. ^ "Aplodontia". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  521. ^ a b c d e f g h i j Nowak, pp. 1627–1635
  522. ^ a b c d e f g h i Chernasky; Motis; Burgin, pp. 402–403
  523. ^ "Glirulus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  524. ^ "Glis". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  525. ^ "Graphiurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  526. ^ "Chaetocauda". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  527. ^ "Dryomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  528. ^ "Eliomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  529. ^ "Muscardinus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  530. ^ "Myomimus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  531. ^ "Selevinia". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  532. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi Nowak, pp. 1248–1306
  533. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh Chernasky; Motis; Burgin, pp. 375–401
  534. ^ "Callosciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  535. ^ "Dremomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  536. ^ "Exilisciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  537. ^ "Funambulus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  538. ^ "Glyphotes". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  539. ^ "Hyosciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  540. ^ "Lariscus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  541. ^ "Menetes". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  542. ^ "Nannosciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  543. ^ "Prosciurillus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  544. ^ "Rhinosciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  545. ^ "Rubrisciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  546. ^ "Sundasciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  547. ^ "Tamiops". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  548. ^ "Ratufa". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  549. ^ "Sciurillus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  550. ^ "Aeretes". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  551. ^ "Aeromys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  552. ^ "Belomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  553. ^ "Biswamoyopterus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  554. ^ "Eoglaucomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  555. ^ "Eupetaurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  556. ^ "Glaucomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  557. ^ "Hylopetes". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  558. ^ "Iomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  559. ^ "Microsciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  560. ^ "Petaurillus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  561. ^ "Petaurista". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  562. ^ "Petinomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  563. ^ "Pteromys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  564. ^ "Pteromyscus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  565. ^ "Rheithrosciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  566. ^ "Sciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  567. ^ "Syntheosciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  568. ^ "Tamiasciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  569. ^ "Trogopterus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  570. ^ "Ammospermophilus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  571. ^ "Atlantoxerus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  572. ^ "Callospermophilus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  573. ^ "Cynomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  574. ^ "Epixerus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  575. ^ "Eutamias". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  576. ^ "Funisciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  577. ^ "Heliosciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  578. ^ "Ictidomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  579. ^ "Marmota". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  580. ^ "Myosciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  581. ^ "Neotamias". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  582. ^ "Notocitellus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  583. ^ "Otospermophilus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  584. ^ "Paraxerus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  585. ^ "Poliocitellus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  586. ^ "Protoxerus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  587. ^ "Sciurotamias". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  588. ^ "Spermophilopsis". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  589. ^ "Spermophilus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  590. ^ "Tamias". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  591. ^ "Urocitellus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  592. ^ "Xerospermophilus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  593. ^ "Xerus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.

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